OOLOGY KAR 1 9 ::?9 The person charging this material is re- sponsible for its return to the library from which it was withdrawn on or before the Latest Date stamped below. Theft, mutilation, and underlining of books ore reasons for disciplinary action and may result in dismissal from the University. To renew call Telephone Center, 333-84OO UNIVERSITY OF ILLINOIS LIBRARY AT URBANA-CHAMPAIGN L161 O-1096 FIELDIANA: BOTANY A Continuation of the BOTANICAL SERIES of FIELD MUSEUM OF NATURAL HISTORY VOLUME 31 The Library of the JAN 1 8 1979 FIELD MUSEUM OF NATURAL HISTORY CHICAGO, U. S. A. 520.5 -F5 vi 31 TABLE OF CONTENTS PAGE 1. Two New Species of Palms from Nicaragua. By S. F. Glassman ... 1 2. Tropical American Plants, VI. By Louis O. Williams 11 3. Agriculture, Tehuacan Valley. By C. Earle Smith, Jr 49 4. Flora, Tehuacan Valley. By C. Earle Smith, Jr 101 5. Preliminary Studies in the Palm Genus Syagrus Mart, and Its Allies. By S. F. Glassman 145 6. Tropical American Plants, VII. By Louis 0. Williams 165 7. Supplement to Orchids of Guatemala. By Donovan S. Correll .... 175 8. Preliminary Notes on Scrophulariaceae of Peru. By Gabriel Edwin . . 223 9. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 233 10. Tropical American Plants, VIII. By Louis O. Williams 247 11. Notes on the Flora of Costa Rica, I. By William C. Burger 273 12. A New Eurystyles from Nicaragua. By Alfonso H. Heller 279 13. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 285 14. A Revision of the Family Geastraceae. By Patricio Ponce de Leon . . 303 15. Studies in American Plants. By Dorothy N. Gibson 353 16. Two New Nicaraguan Juglandaceae. By Antonio Molino R 357 17. Studies in the Palm Genus Syagrus Mart. By S. F. Glassman .... 363 18. Tropical American Plants, IX. By Louis O. Williams 401 --7 PRELIMINARY STUDIES IN THE PALM GENUS SYAGRUS MART. AND ITS ALLIES S. F. GLASSMAN mUMARTIfTHE TROPICAL AMERICAN PLANTS, VII LOUIS 0. WILLIAMS ft waian or ILLINOIS SUPPLEMENT TO ORCHIDS OF GUATEMALA AND BRITISH HONDURAS DONOVAN S. CORRELL I oo CM FIELDIANA: BOTANY VOLUME 31, NUMBERS 5, 6 AND 7 Published by CHICAGO NATURAL HISTORY MUSEUM DECEMBER 22, 1965 BIOLOGY 101 BURRILL HALJJ SUPPLEMENT TO ORCHIDS OF GUATEMALA AND BRITISH HONDURAS DONOVAN S. CORRELL Principal Botanist, Texas Research Foundation FIELDIANA: BOTANY VOLUME 31, NUMBER 7 Published by CHICAGO NATURAL HISTORY MUSEUM DECEMBER 22, 1965 PUBLICATION 999 The author wishes to acknowledge the assistance of Mr. Charles Schweinfurth during the preparation of this paper. Library of Congress Catalog Card Number: 65-291 73 PRINTED IN THE UNITED STATES OF AMERICA BY CHICAGO NATURAL HISTORY MUSEUM PRESS Supplement to the Orchids of Guatemala (and British Honduras) Since the appearance of Oakes Ames' and my work on the orchids of Guatemala (including British Honduras), published by Chicago Natural History Museum (1952-1953, Fieldiana: Botany, 26, Nos. 1 and 2), important collections of several individuals have been made available to us. These are notably those of Percy H. Gentle from British Honduras, and C. L. Lundell and Elias Contreras from the Department of Pete"n, Guatemala, deposited in the Lundell Her- barium, Chicago Natural History Museum, and elsewhere. In addi- tion to these, there are some specimens of various collectors in the United States National Herbarium which were inadvertently omitted in the original publication. In view of the continued, if not increasing, interest in the orchids of this part of Latin America, it was thought advisable to make avail- able this additional information. In citing supplementary distri- bution data for each taxon, page reference is made to where it is described and treated in Orchids of Guatemala. In the case of new additions to the flora, a brief description is included of the taxon, and the salient characteristics that distinguish it from all other presently known orchid taxa in Guatemala and British Honduras are given. Orchids of Guatemala included a total of 527 species or 552 specific and infraspecific taxa in 89 genera. The present publication brings to 542 species or 567 specific and infraspecific taxa in 90 genera the number now known to occur in Guatemala and British Honduras. One new species, Pleurothallis tikalensis, and one new combina- tion, Scaphyglottis major, are proposed. The genus Warrea is defi- nitely new to our flora and apparently the genus Epistephium should be included. The species new to our flora are Habenaria Leprieurii, Vanilla Hartii, Elleanthus caricoides, Spiranthes pseudogoodyerioides, Pleurothallis gelida, P. ovatilabia, Epidendrum cystosum, E. equitanti- folium, E. sculptum, Bletia tenuifolia, Warrea costaricensis, Coryanthes maculata, lonopsis satyrioides and Campylocentrum hondurense. 177 178 FIELDIANA: BOTANY, VOLUME 31 ENUMERATION OF SPECIES Habenaria bractescens Lindl.; Ames & Correll, p. 28, fig. 4. Guatemala. Pete"n: Dos Lagunas, Contreras 1771. Habenaria clypeata Lindl. ; Ames & Correll, p. 28, fig. 5. British Honduras. Toledo: Monkey River, between Monkey River and Cockscomb, Gentle 4229A. Habenaria distans Griseb. ; Ames & Correll, p. 32, fig. 6. Guatemala. Pete"n: Dolores, between kms. 86/87, east of Macha- quila Road, Contreras 2976. Habenaria Leprieurii Reichb. f. Linnaea 19: 376. 1846; Schultes, Native Orchids of Trinidad and Tobago 28. 1960. H. mesodactyla Griseb. Fl. Brit. W. Ind. Isl. 644. 1864; Ames, Orch. 4: 211, pi. 70. 1910; Williams, Ceiba 5(1): 14. 1956. H. Cruegeri Cogn. in Urban, Symb. Ant. 6: 303. 1910. On sandy open pine ridges and uplands. Uncommon in British Honduras, Panama, Trinidad, and northern South America. Plant slender, rigidly erect, up to 5 dm. tall, with one or more tuberoids at base. Leaves linear-setaceous, distant. Flowers greenish yellow, distantly placed in a 1- to 7-flowered spike. Bracts ovate-lanceolate to lanceolate, acuminate, 1- 1.5 cm. long, exceeded by the incurved conspicuously ribbed pedicellate ovary. Sepals often with prominently ciliate margins, 5-6 mm. long; dorsal sepal broadly ovate-elliptic, obtuse, concave, about 3 mm. wide; lateral sepals obliquely ovate- lanceolate, acuminate, about 2 mm. wide below middle. Petals bipartite; anterior lobe setaceous, about 5 mm. long, usually much exceeding the linear posterior lobe. Lip tripartite, about 6 mm. long, the base about 2 mm. wide; lateral lobes linear to setaceous, curved, mostly somewhat exceeding the linear pendulous mid-lobe. This species was not included in Ames and Correll (1952), but Williams (1956) reported it from British Honduras. It is quite pos- sible that our plant should be at least varietally segregated from typ- ical H. Leprieurii. It differs from the island plant in having larger flowers with the lateral lobes of the lip exceeding the mid-lobe and the anterior lobe of the petals exceeding the posterior lobe. Also the sepals are often conspicuously cellular-ciliate, a characteristic not noted in typical H. Leprieurii. The flowers of our plants agree per- fectly with the illustration of H. mesodactyla in Ames, but it does not agree with any of the descriptions published of this plant, including the latest by Schultes. British Honduras. El Cayo: Mountain Pine Ridge, San Augus- tin, Lundell 6712. Stann Creek: Commerce Bight Pine Ridge, Gentle CORRELL: ORCHIDS OF GUATEMALA 179 8291, 8361. Toledo: near Jenkins Creek, north of Monkey River, Gentle 4097. Punta Gorda-San Antonio Road, Gentle 6597. Habenaria novemfida Lindl. ; Ames & Correll, p. 38, fig. 5. British Honduras. Toledo: in sacatal near Machaca, Gentle 6898. Habenaria pauciflora (Lindl.) Reichb. f.; Ames & Correll, p. 39. British Honduras. Toledo: near Jenkins Creek, north of Monkey River, Gentle 4036. Habenaria repens Nutt. ; Ames & Correll, p. 42, figs. 5, 9. Guatemala. Solola: Panajachel, swamp along Lake Atitlan, 0. Degener & I. Degener 558. Pete"n: Tikal National Park, Aguada Terminos, Lundell 16609. British Honduras. Stann Creek: Commerce Bight Pine Ridge, Gentle 8336. Toledo: 12 Miles, Punta Gorda-San Antonio Road, Gentle 6596. Habenaria strictissima var. odontopetala (Reichb. f.) L. Wms.; Ames & Correll, p. 46, figs. 5, 10. Guatemala. Pete"n: Tikal National Park, Bajo de Santa FC", Ar- royo Corriental, on Aguada Terminos Road, Contreras 1934. Vanilla Hartii Rolfe Kew Bull. (1899) 133; Porteres in Bouriquet Le Vanillier et la Vanille (Encycl. Biol. XLVI) 250, fig. 113. 1954; Schultes Native Orchids of Trinidad and Tobago 39. 1960. Climbing on trees in moist forests at low elevations, commonly on ridges. Uncommon in Trinidad, Tobago, British Honduras, Guate- mala, Honduras (Dept. Atlantida, Standley 52824), Costa Rica (Prov. Limon, Standley & Valeria 48905), and Panama (Prov. Panama, Standley 29353). Stem slender. Leaves with a short twisted petiole up to 1 cm. long; blade fleshy but thinnish, elliptic-lanceolate to ovate-lanceolate or oblong-elliptic, more or less abruptly acuminate, up to 11 cm. long and 3.5 cm. wide, usually smaller, about as long as the internodes. Inflorescence axillary, compactly few-flowered, the floral axis 2 cm. or less long. Floral bracts broadly ovate, rarely more than 5 mm. long. Flowers white. Sepals and petals similar, very narrowly linear-oblanceolate, narrowly obtuse at apex, up to about 4 cm. long and 7 cm. wide above the middle. Lip attached to the column almost to its apex to form a tube, with the tube slightly inflated at base, about 4 cm. long from its attachment at base of column to the crenulate-erose apical lamina, about 1.5 cm. wide across the free apical portion when spread out; disk with veins somewhat thickened toward apex of lip, with a small tuft of retrorse centrally placed plates arising about 12 mm. below apex of lip. Fruit short, 5-8 cm. long. 180 FIELDIANA: BOTANY, VOLUME 31 This species, which until now was thought to be endemic to Trini- dad and Tobago, is new to Central America. The Central American plants have, on the whole, larger leaves than those described for the island plants by both Porteres and Schultes, but Schweinfurth and I can find no clear distinction for separating the plants from these two regions. It may be distinguished from all other vanilla-species in our area not only by the floral difference and the long-acuminate leaves but by the extremely abbreviated floral axis. Guatemala. Pete"n: Dolores, on Rio Mopan trail, Contreras 3063. British Honduras. Toledo: Bolo Camp, upper reach, Golden Stream, Gentle 4560. Between Orange Point and Moho River, Gen- tle 7673. Vanilla Pfaviana Reichb. f.; Ames & Correll, p. 55, fig. 15. British Honduras. Toledo: near river beyond Columbia, Gentle 6152. Near ocotal pine ridge, 1-3 miles in trail from 7 miles Punta Gorda-San Antonio Road, Gentle 6957, 7078. Feeders Road leading to Big Fall, Gentle 7023. Near Columbia, Gentle 7108. Vanilla planifolia Andrews; Ames & Correll, p. 57, fig. 16. Guatemala. Pete"n: Tikal National Park, Tikal, Contreras 1841 (narrow-leaved form) . About 2 km. south of Dolores, Contreras 2239. Tikal National Park, Bajo de Santa Fe\ Arroyo Corriental, on Aguada Terminos road, Lundell 15818, 15940. British Honduras. Toledo: Bolo Camp, upper reach of Golden Stream, Gentle 4521. Canada Hill-Alta Visa Road, Gentle 8054. The following collections have their inflorescences terminating a leafy branch or a mainstem. This is not a common occurrence, at least not in herbarium specimens, and it might be of some significance in this species. British Honduras. Toledo: Near Jacinto Hills, Gentle 5106. Feed- ers Road, 14 Miles, San Antonio-Punta Gorda Road, Gentle 6721. Beyond San Antonio, Gentle 7557. Vanilla pompona Schiede; Ames & Correll, p. 59. Guatemala. Pete*n: Dolores, about 800 m. south of the village on the Machaquila Road, Contreras 2333. Dolores, on Rio Mopan trail, Contreras 3063. Canchacan, Lundell 16457. Epistephium sp. An unidentifiable specimen from British Hon- duras which I believe belongs to this genus is in the Ames Orchid CORRELL: ORCHIDS OF GUATEMALA 181 Herbarium at Harvard University. If correct, this would be its only known occurrence in North America. Elleanthus caricoides Nash, Bull. Torr. Bot. Club 34: 119, pi. 7, 1907. Epiphytic on trees in moist forests and groves. Uncommon in British Honduras, Nicaragua and Costa Rica. Plant densely tufted, essentially smooth and glabrous, up to 4 dm. tall. Stems slender, numerous, fasciculate, terete, about 2 mm. in diameter, sparsely spotted with purple. Leaves 5 to 7; sheaths grooved, spotted with purple, much shorter than the blades; blades flat, shining, 7-nerved, erect, somewhat twisted and re- curved above base, linear to linear-lanceolate, long-acuminate at apex, narrowed and sheathing at base, 1-2 dm. long, about 7 mm. wide, reduced above. Inflores- cence congested and subcapitate, 2-3 cm. long, on a peduncle less than 1.5 cm. long. Floral bracts scarious, ovate-lanceolate, long-acuminate, greenish, 2 to 3 times as long as the white flowers, nearly erect and recurved above the middle. Flowers and ovary appressed-pubescent with black-brown hairs, about 1 cm. long, gibbous on one side. Lateral sepals oblong to oblong-elliptic, rather abruptly con- tracted above into a laterally compressed and keeled apex, 7-8 mm. long, about 3 mm. wide. Dorsal sepal 5-6 mm. long and about 2 mm. wide. Petals linear- oblong, somewhat dilated at the rounded and apiculate apex, 6-7 mm. long, 1.5- 2 mm. wide. Lip nearly campanulate, completely surrounding the column so that its margins meet, 7-8 mm. long, when spread out 10-12 mm. wide at the truncate and sparingly ciliate apex, 5-6 mm. wide at the base; disk with a 2-ridged crest just below the middle, the 2 oblong appendages near the base of the lip oblong and 1.5-2 mm. long and 1 mm. wide. Column 7-8 mm. long, 4-toothed at the broadened apex. This species, which was not included in Ames and Correll (1952), is new to British Honduras. It may be distinguished from all other species of Elleanthus (cf . Ames and Correll, p. 60) found in our region, especially E. poiformis Schltr., by its narrowly linear-lanceolate leaves and the tight subcapitate inflorescence with long-attenuate floral bracts that greatly exceed the flowers. British Honduras. Toledo: Between Rancho Chico and Cocks- comb, Monkey River, Gentle 4422. On hill top, Balsam Hill, Edwards Road, beyond Columbia, Gentle 6329. Elleanthus linifolius Presl; Ames & Correll, p. 62. Guatemala. Pete" n: Santo Toribio, north of road, Contreras 2753. Dolores, on Machaquila Road, between kms. 85-87, Contreras 3021, 3135. British Honduras. Stann Creek: Dry Creek, Humming Bird Highway, Gentle 8399. Toledo: Jacinto Creek, Rio Grande, Gentle 4924. Punta Gorda-Seven Hills Road, ascending the Rio Grande, 182 FIELDIANA: BOTANY, VOLUME 31 Gentle 5003. Balsam Hill, Edwards Road, Gentle 6331. On hill slope beyond Pate's Camp, Edwards Road beyond Columbia, Gentle 7159. Sobralia decora Batem.; Ames & Correll, p. 64, figs. 18, 19. Guatemala. Alta Verapaz, Cacao, Finca Trece Aguas, G. P. Goll 48. Pet