PHYSIOLOGY STOMATOGNATHIC SYSTEM
PART-5
CONTENTS
INTRODUCTION
FUNCTIONAL ANATOMY OF ORAL CAVITY
SALIVARY GLANDS
PHYSIOLOGY OF SALIVA SECRETION
MASTICATORY SYSTEM
PAIN PHYSIOLOGY
DEGLUTITION
SPEECH
CONCLUSION
REFERENCES
INTRODUCTION
PHYSIOLOGY
Concerned with the functioning of organism and the processes , function of all or part of an organism
STOMATOGNATHIC SYSTEM
Structures of the mouth and jaws considered collectively as they subserve the function of mastication ,
deglutition ,respiration and speech
FUNCTIONAL ANATOMY OF ORAL CAVITY
Oral cavity also known as buccal cavity. Oral cavity opens anteriorly to the exterior through the lips
and posteriorly through fauces into PHARYNX.digestive juice present in the mouth is saliva which is
secreted by salivary glands
FUNCTIONS OF ORAL CAVITY
1. Ingestion of food materials
2. Chewing of food and mixing with saliva
3. Appreciation of taste of the food
4. Deglutition
5. Speech
6. Smiling and other expressions
SALIVARY GLANDS
SALIVA
Saliva is the clear mucoserous secretion formed mainly in salivary glands and has lubricative,
cleansing & anti microbial, excretory and digestive functions.
Oral fluid also known as whole saliva or total saliva is composed primarily of saliva secreted by
major and minor salivary glands.
Classification of salivary glands
Based on size and location
MAJOR SALIVARY GLAND MINOR SALIVARY GLANDS
Parotid Labial & buccal glands
Submandibular Palatine glands
Sublingual Glossopalatine glands
Anterior lingual/Glands
of
Blandin & Nuhn
Posterior lingual
Glands of Von Ebner
BASED ON THE HISTOCHEMICAL NATURE OF THE SECRETORY PRODUCT
MUCOUS
Labial & buccal glands
Glossopalatine
Palatine
Anterior lingual
SEROUS
Parotid
Glands of von Ebner
MIXED
Submandibular (serous predominant)
Sublingual (mucous predominant)
Based on the duct system
SIMPLE Minor Salivary Glands
COMPOUND Submandibular
Parotid
Sublingual
STRUCTURE OF THE SALIVARY GLAND
The working parts of the salivary glandular tissue
consists of secretory end pieces (acini),
myoepithelial cells and the branched ductal
system.
Serous glands – roughly spherical
Mucous glands – tubular configuration
DUCTAL SYSTEM :
1.Intercalated duct
2.Striated duct
3.Excretory duct
NERVE SUPPLY OF SALIVARY GLANDS
-Salivary glands are purely under nervous control .
-Nerve supply is from the sympathetic and parasympathetic branches of autonomic nervous
system.
-Stimulation of sympathetic supply – release of amylase and vasoconstrictor
-Parasympathetic nerves are secretomotor
-These nerves innervate the acinar cells,duct cells,blood vessels and myoepithelial cells.
FORMATION OF SALIVA
TWO STAGE HYPOTHESIS
OF SALIVA FORMATION
PROPERTIES OF SALIVA
• VOLUME-1000-1500 ML/DAYPH
• PH-6.35-6.85
• TONICITY-SALIVA IS HYPOTONIC TO PLASMA
• SPECIFIC GRAVITY-1.002-1.12
Applied Aspects
The main causes of hyposalivation are
1) Ageing
2) Iatrogenic
Drugs
Irradiation
1) Disease
• Salivary gland disease
Salivary aplasia (agenesis)
Sjogren’s syndrome
Infections
1) Dehydration
2) Psychogenic (anxiety, stress)
SALIVARY GLAND HYPERFUNCTION
Drugs
-cevimiline
-physostigmine
-pilocarpine
Other conditions
-parkinson’s disease
-heavy metal poisoning
Composition of saliva
PHYSIOLOGY OF GUSTATION
TRANSMISSION OF TASTE SIGNALS TO CN
TASTE PAPILLAE
TASTE BUDS
Taste buds are group of 30-100 individual elongated neuroepithelial cells
Which are 50-60 microns in height and 30-70 microns in width which are embedded in
epithelium
MECHANISM OF STIMULATION OF TASTE BUD
Stimulating substance or tastant
Binding to microvilli ( protein receptor molecule)
Stimulates exocytosis and release of neurotransmitter from vesicles
Opening of ion channels and entry of Na ions
Depolarisation of cell membrane
PRIMARY SENSES OF TASTE
SALIVA-TASTE INTERACTION
Gustatory function at the peripheral level requires 3 factors-
A molecule that elicits taste
A taste receptor cell
An aqueous environment provided by saliva
Saliva contributes the tasting ability by-
First as a solvent of food
Second as a carrier of taste eliciting molecules
Third through its composition
Hence when the salivary output is affected adversely , taste function can concomitantly affected
TASTE DYSFUNCTION
Loss or impairment of taste can occur in degrees
NORMOGEUSIA
HYPOGEUSIA
DYSGEUSIA
ALIAGEUSIA
PHANTOGEUSIA
AGEUSIA
APPLIED ASPECTS
MASTICATORY SYSTEM
Masticatory system includes
-Hard tissue components
-Soft tissue components
STRUCTURE OF SKELETAL MUSCLE
MUSCLE FIBER
The end of the muscle fiber fuses with a tendon fiber
Tendon fibers in turn collect into bundles to form the muscle tendon that inserts into the bone
Muscle fiber contains several hundred to several thousand myofibrils
Each myofibril in turn lying side by side, about 1500 myosin filaments and 3000 actin filaments, which
are large polymerized protein molecules that are responsible for muscle contraction.
CLASSIFICATION OF MUSCLE FIBRES
Muscle fibers can be characterized by type according to the amount myoglobin
SLOW MUSLE FIBRES OR TYPE I MUSLE FIBERS OR RED MUSCLES
FAST MUSLE FIBERS OR TYPE 2 MUSLE FIBERS OR PALE MUSCLES
Based on functional and histological appearance
TYPE S or Slow , fatigue resistant fibres
TYPE FR or fatigue resistant fibres
TYPE FF or fast fatigable fibres
TYPES OF MUSCLE CONTRACTION
ISOTONIC CONTRACTION
ISOMETRIC CONTRACTION
MUSCLES OF MASTICATION
FUNCTIONS OF MASSETER MUSCLE
As fibers of the masseter contract, the mandible is elevated and the teeth are brought into contact.
Its superficial portion may also aid in protruding the mandible.
When the mandible is protruded and biting force is applied, the fibers of the deep portion stabilize the
condyle against the articular eminence.
FUNCTION OF TEMPORALIS MUSCLE
When the temporal muscle contracts, it elevates the mandible and the teeth are brought into contact.
When the anterior portion contracts, the mandible is raised vertically
Contraction of the middle portion will elevate and retrude the mandible.
Posterior fibers below the root of the zygomatic process contraction will cause elevation and slight
retrusion.
FUNCTION OF MEDIAL PTERYGOID
Along with the masseter, it forms a muscular sling that supports the mandible at the mandibular
angle.
When its fibers contract, the mandible is elevated and the teeth are brought into contact.
This muscle is also active in protruding the mandible
Unilateral contraction will bring about a mediotrusive movement of the mandible.
JAW OPENING MUSCLES
FUNCTIONS OF INFERIOR LATERAL PTERYGOID
When the right and left inferior lateral pterygoids contracts simultaneously
condyles are pulled down the articular eminences and the mandible is protruded.
Unilateral contraction creates a mediotrusive movement of that condyle and causes a lateral movement
of the mandible to the opposite side.
When this muscle functions with the mandibular depressors,the mandible is lowered and the condyles
glide forward and downward on the articular eminence
Functions of superior lateral pterygoid
Although the inferior lateral pterygoid is active during opening, the superior remains inactive .Becoming
active only in conjunction with the elevator muscles.
The superior lateral pterygoid is especially active during the power stroke and when the teeth are held
together.
The powerstroke refers to movements that involve closure of the mandible against resistance, such as in
chewing or clenching the teeth together.
DIGASTRIC MUSCLE
FUNCTION-elevates the hyoid and depresses mandible
Mylohyoid muscle
FUNCTION-elevates hyoid and floor of the mouth
and Depresses the mandible
FUNCTION OF BUCCINATOR MUSCLE
Purpose is to pull back the angle of the mouth and to flatten the cheek area –which aids in
holding the cheek during chewing
This action causes the muscle to push back the food on the occlusal surface of posterior teeth
It aids whistling and smiling
In neonates it is used to suckle
CONTROL OF MANDIBULAR REST POSITION
THE HEAD IS STATIONARY - VISCO –ELASTIC FORCES
HEAD IS MOVING VIGOROUSLY UP AND DOWN DURING RUNNING AND JUMPING STRETCH
REFLEXES
• CHEWING CYCLE
• Preparatory phase
• Food contact phase
• Food crushing phase
• Tooth contact phase
MECHANISMS THAT MODULATE JAW MUSCLE ACTIVITY DURING CHEWING
MUSCLE SPINDLE RECEPTORS
MECHANORECEPTORS
BITE FORCES DURING NORMAL MASTICATION
Maximum biting force is upto 350 N
When chewing western diets,jaw closing muscles exert only 30-40% of their maximum force
The maximum bite force of who have lost all their natural is decreased by 50% ,even when they
wear well fitting dentures
MASTICATORY MUSCLE VS NORMAL MUSCLES
The extraordinary force exerted by masticatory muscles is due to-
fiber composition
Biomechanics
Most muscles act via long tendons and insert onto bones - poor mechanical advantage
Eg-elbows ,knees
Where jaw muscles act directly across the temporomandibular joint - good mechanical
advantage and with no need of tendon
APPLIED ASPECTS
MUSCLE PATHOLOGY
MUSCULAR DYSTROPHY –
DUCHENNE’S MUSCULAR DYSTROPHY
-Becker muscular dystrophy
MYOSITIS - inflammation of the muscle that results from local causes such as traumatic injuries,muscular
strain,orodental infections
Myofascial pain syndrome - it is a pain disorder in which unilateral pain is referred from the trigger
points in myofascial stuctures to the muscles of the head and neck
Causes-
sudden mouth opening
Phyisiological stress
Bad posture
Muscular tension
Bruxism
NEUROMUSCULAR JUNCTION DISEASES
-MYASTHENIA GRAVIS
-LAMBERT-EATON SYNDROME
DEGLUTITION
DEGLUTITION
-The process or act of swallowing which is carried by stomatognathic system
Phases of deglutition –
INFANTILE SWALLOW
MATURATION SWALLOW
Oral preparatory phase
Pharyngeal phase
Oesophageal phase
INFANTILE SWALLOW
Swallowing pattern is purely reflex in nature in newborn(involuntary)
The ability to feed from the breast is present in the newborn child by negative pressure
The milk is directed continuously to the pharynx by an autonomous peristalitic movement of
tongue and mylohyoid muscle
During swallowing-
the entrance to infant larynx is elevated
Then epiglottis placed behind the soft palate
Which is then possitioned in the vallecular space just infront of the epiglottis
Functionally continues tube from the nose through the larynx to the bronchi
This enables uninterrupted breathing even during swallowing
DISTINGUISING FEATURES BETWEEN MATURE AND INFANTILE SWALLOW
The jaws are apart and the tongue is placed between the upper and lower gum pads
The mandible is stabilised by the contraction of the muscles of facial expression(innervated by
facial nerve) and interposed tongue
The swallow is guided and to large extent controlled by the sensory interchange between lips
and tongue
Mature swallow
Oral preparatory phase
It is voluntary stage
The anterior of the tongue is raised and pressed against the hard palate by superior
longitudinal and transverse muscles
The movement takesplace from anterior to posterior side
This pushes the food bolus into the posterior part
The soft palate closes down on to the back of the tongue and helps to form bolus
Next the hyoid bone is moved upwards and forwards by the suprahyoid muscles
The posterior part is supported by styloglossi
The palatoglossal arches are approximated by palatoglossi
This pushes the bolus through the oropharyngeal isthmus to the oropharynx
Now next stage begins
PHARYNGEAL PHASE
It is an involuntary stage
During this stage food is pushed from the oropharynx to the lower part of the laryngopharynx
Nasopharyngeal isthmus is closed by the soft palate by levator veli palatani and tenser veli palatani
This prevents the food bolus from entering the nose
PHARYNGEAL MUSCLE CONTRACTION:
Soft palate is pulled upward and prevents the reflux of food to nasal cavity
Palatopharyngeal folds are pulled medially to approximate each other – form a saggital slit
Vocal cords are approximated
Larynx is pulled upward & anterior by neck muscles
Epiglottis swing backward over the opening of larynx
Upward movement of larynx & enlargement the opening of esophagus
Upper 3-4cm of esophagus relaxes
Muscular wall of pharynx contracts to push the food downward (propulsive contraction)
OESOPHAGEAL PHASE
This phage commences as soon as the food passes the cricopharyngeal sphincter
Peristalitic activity of the oesophageal walls occur to pass the food into the stomach
Tongue and palate return to their original position to start the next cycle
ESOPHAGEAL STAGE OF SWALLOWING
Conducts food rapidly to the stomach
Two types of peristaltic movements:
1° peristalsis:
– continuation of a peristaltic wave
– begins in pharynx & spreads into esophagus
– passes in 8-10 sec
2° peristaltic waves:
– results from the distention of esophagus
– begins if the 1° wave failed to push the food down
Esophageal reflux can be prevented by
Gastro-esophageal sphincter
Valve-like mechanism: short portion of the esophagus that extends beneath the diaphragm
before opening into stomach
Applied aspects
BULIMIA NERVOSA
ANOREXIA NERVOSA
GASTRO-OESOPHAGEAL REFLEX DISEASE
PLUMMER-VINSON SYNDROME
Speech vs language
LANGUAGE
Language comprises the central function associated with Processing of linguistic information in the brain
SPEECH AND HEARING
It denotes the peripheral processes that are needed to produce spoken language and to receive spoken
utterances
SLEEP APNEA
Central sleep apnea - which occurs when the brain fails to send important signals to the breathing
muscles
Obstructive sleep apnea - which occurs when air cannot flow through the nose or mouth even though
the body is still trying to breathe.
Obstructive sleep apnea is far more prevalent and easily treatable by the dentist.
This disorder causes significant morbidity, particularly in terms of daytime functioning and its impact on
quality of life.
Upper airway space
REFERENCES
Clinical oral physiology- timothy et al
Clinical oral sciences– malcolm harris, edgar and meghji
Human anatomy volume -3 b.d chaurasia th edition
Atlas of anatomy-netter 3rd edition
Grays anatomy
Human physiology-a.k.jain
GUYTON-PHYSIOLOGY- 11 TH EDITION
OKESON - MANAGEMENT OF TEMPOROMANDIBULAR JOINT DISORDERS AND OCCLUSION -
6TH EDITION
Salivary diagnostics , DAVID T.WONG