Module 3
Module 3
Introduction
       Biological diversity or biodiversity refers to the variety and abundance of different species of
        plants and animals
            o Biodiversity can be affected slowly or quickly over time by natural selection
                pressures
            o Human impact can affect biodiversity over a shorter time period
            o The general health of the planet depends on biodiversity. In all ecosystems, the
                greater the biodiversity the more stable and resilient the system becomes
Organising the ecosystem
Ecosystem: made up of organisms (biotic) living in an area that interact with each other and with the
non-living (abiotic) environment in which they live
Community: groups of different species living in the same habitat and interacting with one another
Population: group of organisms of the same species living together in a defined geographical area
Organism: an individual of a species
3.1.1 predict the effects of selection pressures on organisms in ecosystems, including:
       Biotic factors
       Abiotic factors
Selection pressures
       Selection pressures are changes in the environment that put constraints on organisms and
        determine which individuals are best suited to the prevailing conditions
       Abiotic factors or non-living environment (e.g. amount of water, temperature, amount of
        light, types of rocks and soil etc.) and how these affect the biotic environment
       Biotic factors or living organisms (plants, animals, microbes etc.) that live in the area and
        how these living things interact with each other
Abundance and distribution
Distribution: of a species describes where it is found
Abundance: of a species determines how many individuals of that species live throughout the
ecosystem
        Organisms occupy the areas where the biotic and abiotic factors of the environments suit
         them. They live where their chances of survival are high, where their requirements for
         survival are met, and where they are able to avoid predators
        Abiotic and biotic selections pressures affect the distribution and abundance of organisms in
         an ecosystem by causing fluctuations or changes in population numbers and movement.
         Populations may occupy certain areas and not others due to the resources that are available
Abiotic factors
       Abiotic factors are the non-living factors in an environment including:
           o Temperature
           o Water availability
           o Air pressure
             o   Gas (O2 and CO2) concentration
             o   Light intensity
             o   Salinity
Biotic factors
       Biotic factors are the living factors in an environment including:
            o Availability and abundance of food
            o Number of competitors
            o Number of mates
            o Number of predators
            o Number and variety of disease-causing organisms
      Behavioural adaptations
Behavioural adaptations
      Behavioural adaptations are actions performed by an organism in response to a stimulus
       that improves its chances of survival
      For example: birds migrating to avoid extreme temperatures
Behavioural Adaptations of Mangrove Crabs
      Mangrove crabs burrow sideways into the soft mud to gain protection from both dehydration
       and predators. They use the water in their burrows to keep their gills moist and keep away
       from the hot sun
Movement Adaptations of Plants
      Plants generally do not move much, but they still do exhibit responses to stimuli. It is just
       slow and subtle
      For example:
           o sunflowers exhibit phototropism moving in response to sunlight and plant hormones
           o The shy plant Mimosa pudica can change turgor pressure rapidly, collapsing leaves
               inwards when touched by an insect or predator. This protects the plant from harm,
               and is called nastic movement
Nastic Adaptations of Carnivorous Plants
      Venus fly traps and sundews grow in nutrient poor soil, especially lacking nitrates
      When trigger hairs are touched, these plants collapse leaves inwards, catching insects use
       nastic movement
      The plants secrete enzymes to digest insects and release nitrates as liquid fertilizer for the
       plants
3.2.2 investigate, through secondary sources, the observations and collection of data that were
obtained by Charles Darwin to support the Theory of Evolution by Natural Selection, for example:
      Finches of the Galapagos Islands
Observations of Charles Darwin
      The British naturalist Charles Darwin (1809-1882) suggested a persuasive argument to
       explain the range of adaptations and biodiversity on Earth
      His ideas built on previous theories of evolution and was called the Theory of Evolution by
       Natural Selection
Finches of the Galapagos Islands
      Darwin observed that the Galapagos Island finches had beaks adapted for specific types of
       food
      Darwin reasoned that each of the finches evolved from a common ancestor
      He proposed that distinctive adaptations such as beak structure evolved in response to the
       selection pressure of different food sources
      Australian flora and fauna
                              Part 3: Theory of Evolution by Natural Selection
3..3.1 explain biological diversity in terms of the Theory of Evolution by Natural Selection by
examining the changes in and diversification of life since it first appeared on the Earth
Biodiversity
       Biodiversity is the variety of all forms of life on Earth, the diversity of the characteristics that
        living organisms have and the variety of ecosystems of which they are components
             o Genetic diversity is the total amount of genetic characteristics in a species
             o Species diversity is the total number of species in a community
             o Ecosystem diversity is the variation of different ecosystems in an area
Evolution
       Evolution is the change in living organisms overtime
       Recipe for evolution: variation, selection and time
Evolution by Natural Selection
       Charles Darwin and Alfred Wallace jointly proposed a mechanism for species change
       They did not work together but arrived at the same conclusion:
           o Individuals within populations naturally possess variations in structure, behaviour
                and/or functioning
           o If these variations confer an advantage, organisms survive a change in the
                environment. These organisms are able to reproduce and pass on their favourable
                characteristics to future generations (later described as survival of the fittest)
Darwin’s Big Idea
       Darwin’s theory of Evolution by Natural Selection and Isolation has 4 main points:
           o Variation
                    In any population, there is variation within the species
                    Variation comes about due to genetic changes or mutations; these are not
                       changes the individual organism has control over
           o Natural selection
                    Some organisms have variations in their features that make them better suited
                       than others to a changed environment
                    If a population consists of a diverse range of individuals, then the populations
                       is better able to survive a sudden change in the environment
                    This diversity allows some organisms to compete more successfully, and
                       survive and breed and therefore pass on their genes to the next generation
                    This is, those individuals that compete successfully in the new environment
                       outlive those that do not have such variations, this is termed natural
                       selection
           o Survival and reproduction
                    In any population, there are offspring that do not reach maturity and
                       reproduce, possibly because they are not as well suited to their environment,
                       struggle to find adequate nutrition, are more vulnerable to predators etc, and
                       therefore their characteristics are not pass on
           o Isolation
                    If a population is isolated from the original population, individuals that have
                       variations that allow them to survive the changed conditions will reproduce
                       and pass on these characteristics
                      Eventually the population becomes so different to the original and individuals
                       are no longer able to interbreed and produce fertile offspring – a new species
                       has evolved
3.3.2 analyse how an accumulation of microevolutionary changes can drive evolutionary changes and
speciation over time, for example:
      Evolution of the horse
Macro and microevolution
      Evolution can be considered over very long periods of time and over shorter periods of time
          o Macroevolution takes place over millions of years and results in the arising of new
               species
          o Microevolution takes place over a shorter periods of time and results in changes
               within populations, but it does not produce new species in the short term
                    e.g. pepper moths – variation in the species
Microevolution over time leads to speciation
      microevolution involves changes in characteristics over short periods of time
      Mutation and natural selection are the main processes that drive microevolutionary change
      An accumulation of these changes over time can lead to speciation
          o An example of microevolution is the modern day horse
                   Through fossil evidence we know that it has developed from a small dog-
                       sized, forest-dwelling animal called Hyacotherium to the horse of today
Horse Fossils
      Modern horses are large animals with only one toe and a large cheek span
      There are fossils of many transitional forms – fossilised remains of horses with three toes and
       intermediate cheek span
      Early horses were small animals with four toes, short legs and a narrow cheek span between
       the front teeth and cheek teeth
Horse Teeth
      Early horses were browsers and ate fruit. They had cheek teeth capped with enamel very
       similar to humans and other mammals
      Modern horses are grazers and eat grass. They have grinding cheek teeth that grow
       continually throughout the horse’s life
Microevolution of the Horse
      Towards the end of the Oligocene, Earth started cooling and drying. Rainforest gradually
       gave way to grassland
      This produced selective pressures for horse ancestors: change to grass for food, and less
       bushes, undergrowth and shadows for hiding places. Running faster became a better defence
       against predators
      Evolution of the platypus
Mammals, Marsupials and Monotremes
      Mammals are animals with hair or fur, that split into roughly 3 groups:
       1. Placental mammals carry offspring in the uterus
          e.g. humans and cats
        2. Marsupial mammals carry offspring in pouches
           e.g. koala and kangaroo
        3. Monotremes which lay eggs
           e.g. echidna and platypus
Platypus Ancestors
       There are only a few specimens of platypus fossils
       Obdurodon dicksoni was a large, spoon-billed platypus
       Its skull is one of the most perfect fossils know from Riversleigh
       They probably fed on insect larvae, yabbies and other crustaceans, and perhaps small
        vertebrate animals such as frogs and fish
       A closely related species monotrematum sudamericanum
       The animal has been reconstructed from fossil teeth
       Unlike the living platypus these fossil platypuses had functional molar teeth
3.3.3 explain, using examples, how Darwin and Wallace’s Theory of Evolution by Natural Selection
accounts for:
       Convergent evolution
       Divergent evolution
Darwin-Wallace theory
       Darwin and Wallace studied a large numbers of living organisms and observed that
        similarities in structure were common. These similarities could be accounted for in one of two
        ways:
        1. Divergent evolution: the process by which organisms that are related look different due
            to exposure of different selection pressures
        2. Convergent evolution: the process by which organisms that do not have a common
            ancestor develop similar features in response to similar selection pressures in their
            environments
Divergent Evolution
Examples:
       Darwin’s finches diverge from a common body plan. Different selection pressure therefore
        favoured some forms over others
       The rise of the variety of horse species is also typical
Convergent Evolution
       Placental mammals from North America and Europe show similarities to marsupials in
        Australia
       The organisms have been exposed to similar selective pressures and have adaptations to
        similar niches
            o E.g. the echidna and hedgehog both have prickles and eat insects
3.3.4 Explain how punctuated equilibrium is different from the gradual process of natural selection
Darwin’s Theory
       Darwin’s theory of evolution by natural selection proposes that populations change slowly
        and gradually over time.
       Therefore, in the fossil record we should see a gradual change, for any one group of organism,
        from ancestor to descendent
       However, we have very few instances
            o One example, is the evolution of the horse which shows a gradual change over 40
               million years
Punctuated equilibrium
       The theory of punctuated equilibrium proposes that evolution occurs in short bursts of rapid
        change, followed by long periods of stability within populations
       This theory was put forward based on fossil evidence: if evolutionary change is gradual, it
        could be predicted that there would be fossilised remains showing these ongoing changes
       However, many fossilised remains how millions of years going without any noticeable
        evolutionary change to most species
            o E.g. Horseshoe crabs have remained almost unchanged for 445 million years
       2. DNA-DNA hybridisation
      DNA-DNA hybridisation is based on the assumption that DNA
       molecules of closely related species have a similar nucleotide
       base order
      The process involves applying heat to a double stranded DNA
       molecule which causes the complementary strands to separate
      One strand from one species is then combined with on strand
       from another species to form a ‘hybrid’ DNA molecule. The
       more closely matched the base pairs are, the strong the binding
       of the strands
      Heat is once again applied, this time to determine how strongly
       the bases have combined; higher temperatures are required to
       separate hybrid strands that are more strongly combined
      Closely related species have a very similar order of nucleotide bases and so their DNA strands
       combine more strongly than species that are distantly related
Comparative anatomy
      Comparative anatomy is the study of similarities and differences in structure (anatomy) in
       different species of living organisms to determine their evolutionary relatedness
      more similarities in the structure implies that they must be separated from a common ancestor
       more recently
Evidence of divergent evolution
      in organisms that are being compared, similarities in structure suggest descent from a
       common ancestor, whereas differences in structure represent modifications – how organisms
       have evolved to become different.
      This is typical of divergent evolution and the similarities are best explained by common
       descent – that is, sharing a common ancestor
Pentadactyl limb
      The pentadactyl (five-digit) limbs of all vertebrates have the same basic bone plan, but show
       modifications because they are used in different ways. This is an example of a homologous
       feature
Evidence of convergent evolution
      Structures that differ greatly in their basic plan, are said to be analogous. They have started
       off being very different and then have evolved independently to become similar, because they
       were selected to be used for a similar purpose
      The presence of analogous features does not provide evidence for evolutionary relatedness,
       but rather for evolution of structures to serve a common purpose in a common environment
           o Example: the Australian echidna and European hedgehog have both developed
               protective spines to discourage predation but in terms of most other structures, they
               are dissimilar
Vestigial structures
      Vestigial structures are evolutionary remnants of body parts that no longer serve a useful
       function within that population. The presence of a vestigial structure provides evidence of
       common ancestry
           o Example: the presence of a reduced tail (coccyx) in humans and the pelvic bones in
               snakes, are difficult to explain unless they are structures that have become reduced
               because they no longer carry out a useful function in that animal’s lifestyle
Comparative embryology
      Comparative embryology is the comparison of embryos of different species at early stages
       of development
      Embryonic comparisons show that general features of groups of organisms appear early in
       development. More specialised features, which distinguish the members of a group, appear
       later in development
Vertebrate embryos
      Fish, amphibians, birds and mammals all show the presence of gill slits, and tails in
       embryonic development
      The presence of gill slits suggest that they all descended from a common ancestor that lived in
       an aquatic environment
      These slits develop into internal gills in fish, external gills in amphibians and Eustachian
       tubes in mammals
Biogeography
       Biogeography is the study of the geographical distribution of organisms, both living and
        extinct, across the world
       Darwin-Wallace theory of evolution proposes that, for a new species to arise, a group of
        individuals must become isolated. If this is true, the new species should resemble species with
        which they shared a habitat
            o Australia’s unique mammals and flowering plants are believed to have arisen
                 because of isolation of the continent. Australian organisms show similarities to
                 fossils on other southern continents, evidence that they may have had a common
                 origin and later evolved
Flightless birds (ratites)
       Some distribution patterns are best explained by continental drift theory
           o E.g. the present-day distribution of flightless birds suggests that these birds
               originated from a common ancestor on Gondwana and that the different populations
               evolved on the isolated continents as they drifted apart
       Techniques used to date fossils and the evidence produced
Fossil Evidence
       Palaeontology is the study of ancient life preserved in rocks (fossils)
       the fossil record is a catalogue of the occurrence and evolution of living organisms through
        geological time
       unfortunately, the chance of an organism fossilising is small, therefore the fossil record is
        incomplete
Relative dating
       relative dating relies on the assumption that fossils found higher in rock strata are younger
        than the lower fossils, so fossils are dated relative to one another (actual age is not
        determined). Techniques used:
            o stratigraphy: relies on sedimentary rocks being formed in layers with the oldest
                rocks at the bottom and the youngest at the top. Therefore, fossils contained in these
                rocks would display the same trend
              o   biostratigraphy: compares fossils in different strata. Index fossils are useful in
                  determining the rock strata in which they are found. The occurrence of a fossil within
                  two different rock locations indicates that the rock containing the fossil specimens
                  were deposited at about the same time
Absolute dating (radiometric dating)
      absolute dating enables the actual age of the specimen to be determined by measuring
       proportions of naturally occurring radioactive elements (radioisotopes) that are present in the
       specimen
           o each radioisotope decays at a known fixed rate, called its half-life (time taken for half
               the atoms to decay)
           o if the half-life is plotted onto a graph, the decay forms an exponential curve
           o the atom that undergoes radioactive decay is known as the parent nuclide and the
               atom which is formed is known as the daughter nuclide
Radiocarbon dating
      carbon becomes radioactive in the atmosphere; the carbon atom, which normally has 6
       neutrons (carbon-12), gains 2 neutrons (carbon-14)
      carbon-14 is unstable and therefore decays back to nitrogen-14 at a known rate
      since carbon-12 is stable, the amount will stay the same
      the proportion of C-14 to c-12 in a fossil tells us how long ago the plant or animal was alive
3.4.2 explain modern-day examples that demonstrate evolutionary change, for example:
      the cane toad
      antibiotic resistant strains of bacteria
Antibiotics
      antibiotics are chemicals that are able to inhibit the growth of bacteria or can destroy them.
       They target the cell wall and inhibit bacterial metabolism
      the first antibiotic, penicillin has highly successful in treating many bacterial infections.
       However, with the widespread use of antibiotics and the misuse and overuse, bacteria have
       evolved strains that are resistant to many, if not all, of the antibiotics available today
Bacterial Reproduction
      bacteria reproduce by binary fission
      when conditions are favourable they can multiply rapidly with many generations reproducing
       within a single da. Numbers of bacteria quickly enter the millions
      bacteria can also reproduce sexually through conjugation, in which they exchange genetic
       material. This process can lead to variation in a population of bacteria
Antibiotic Resistant Strains
      when exposed to an antibiotic (the selection pressure), the bacteria that have variations that
       are best suited to that environment (resistant to the antibiotic) will survive and reproduce
      therefore, the population will become more resistant to the antibiotics over generations
Example: Golden Staph (MRSA)
      Staphylococcus aureus (golden staph) is a bacterium that causes mostly mild skin infections,
       but can cause serious har, when it infects the blood or lungs
      It has now evolved a strain called MRSA (methicillin-resistant S. aureus) that is resistant to
       many of the antibiotics used to treat it
      Today many people are wary of hospitals de to the rapid rise in resistant bacteria. There are
       many documented cases where a person has entered hospital for a minor infection and has
       caught golden staph