Biological Classification         19
Life Cycles of Some Fungi
These can be described as follows
(i) Life Cycle of Rhizopus
The structural representation (sexual and asexual) of life cycle of
Rhizopus is as follows
                               Fragmentation
                                Vegetative
                                Reproduction
                                 Sporangium
                               Chlamydospore
                                    Oidia
                                  Asexual
                                Reproduction
                                  Rhizopus
                                  mycelia
                                   Sexual              Progametangium(–)
            Germ spores         Reproduction
                   (+ or –)                    Progametangium
                                                      (+)
                                                                Gametangium (–)
       Germ sporangium
                                                  Gametangium
                           Haplophase (n)              (+)
                                                                    Coenogamete (–)
         Promycelium
                                                        Coenogamete
                                                                +
                                 Azygospore
                 Meiosis                                  Fertilisation
                                  Diplophase (2n)
                                     Zygospore
                              Life cycle of Rhizopus
20     Handbook of Biology
(ii) Life Cycle of Yeast
The diagrammatic representation of sexual cycle of Saccharomyces
cerevisiae is as follows
       Dwarf strain
       yeast cells                    B
                                                 Budding                           +
                                                                 C          +
                        A
                                                                         Gametangia
                                                                –
            Germinate
                                                      –
                                               Plasmogamy
                L            Ascospore
                                                                                D
                                                                Karyogamy
                            Ascospores
                                                 e
                                             has
                                          op
                             Mature
                                               )
                                         pl
                                                        e
                                              (n
                                                 (2 has
                             ascus                              Zygote
                                       Ha
                                                      p
                                                   lo
                                                   n)
                                                                                    E
                                                 p
                                              Di
       K                    Young
                                                                          Germination
                            ascus
                                                 Large strain
                                                 yeast cell
                                    Ascus mother                                       F
        J
                                    cell
      Meiosis                                                                          Bud
                                                            G            Budding
                        I
                                      H
                      Life cycle of Saccharomyces cerevisiae
Heterothallism
The phenomenon of having two genetically different and compatible
sexual strains in two different thalli is called heterothallism. It was
discovered by Blakeslee in Mucor.
                                                        Biological Classification   21
Mushroom and Fairy Rings
Agaricus compestris is an edible mushroom. It is also called white
button mushroom. The fruiting body of Agaricus, arises in concentric
rings (called fairy rings or fungal flowers) from the mycelium present
in the soil.
Lichens
They have composite structure and consist of two dissimilar organisms
forming a symbiotic relationship between them.
Lichens are formed by
●  Algal Part — Phycobiont — Provide food to fungi
●  Fungal part — Mycobiont — Provide shelter to algae
Lichens are of three types on the basis of their structure
    (i) Crustose lichens These are point-like, flat lichens, e.g., Caloplaca.
 (ii) Foliose lichens These lichens have leafy structure,
      e.g., Hypogymnia physodes.
(iii) Fruticose lichens These are branched lichen, form
      filamentous branching, e.g., Cladonia evansii, Usnea australis,
      etc.
Various forms of lichens are given below
                      Fungal fructification
                                                          Parmella
                    Graphis                 Cora
                                                          (foliose)
                    (crustose)              (foliose)
                                 Attaching disc
                                   Pendent
                                   branches
                                          Podetia
                                             Early
                                             foliose
                                             part          Cladonia
                                       Fungal              (fruticose)
                   Usnea               fructification
                   (fruticose)
                                      Forms of lichens
Mycorrhiza
It is a symbiotic association between a fungus and a plant. Plants
prepare organic food and supply them to fungus and in return, fungus
supplies water and mineral nutrients to plants.
22       Handbook of Biology
D. Kingdom–Plantae (Eukaryotic, Chlorophyllous Organisms)
These are chlorophyllous and embryo forming organisms. Mostly
non-motile and function as the producers in ecosystem as they can fix
solar energy into chemical energy through the process of
photosynthesis. The cell wall in plants is cellulosic and stored food
material is in the form of starch.
A detailed account of plant kingdom is given in chapter 6.
E. Kingdom–Animalia (Multicellular, Eukaryotic Organisms)
The heterotrophic, eukaryotic organisms which are multicellular and
lack cell wall, present in this kingdom. Animals have advanced level of
tissue organisation, in which the division of labour is highly specific.
The two main groups among animals are Non-chordata and Chordata,
divided on the basis of the presence of notochord in them.
A detailed account of animal kingdom is given in chapter 7.
Viruses and Viroids
1. Viruses
The term ‘Virus’ means poisonous fluid. The word was coined by
Louis Pasteur. Viruses are very small (0.05-0.2 µm), infective,
nucleoprotein particles, which can be called as living because of the
presence of nucleic acid as genetic material and ability to produce their
own copy-viruses. They show only some properties of living beings,
otherwise they behave like non-livings. Hence, these are referred to as
the connecting link between living and non-living.
On the basis of nature of genetic material, the viruses are of two types
   (i) Adenovirus DNA containing, e.g., HIV, etc.
  (ii) Retrovirus RNA containing, e.g., Rous sarcoma virus, etc.
On the basis of their host, the viruses can be categorised as
   (i) Animal virus (Zoophagineae), e.g., HIV, sarcoma, etc.
  (ii) Plant virus (Phytophagineae), e.g., TMV, etc.
 (iii) Bacterial viruses (Phagineae), e.g., T4 phage, etc.
Characteristics of Viruses
Characteristics of viruses are as follows
Living
●
    They can replicate.
●
    In host body, they can synthesise protein.
●
    They cause diseases like other living organisms.
●
    Similar gene mutation as living organism.
                                                  Biological Classification       23
Non-living
●   Do not have protoplasm, and do not perform metabolism.
●   These can be crystallised.
●   They do not respire.
●   In vitro culture is not possible.
Structure of Viruses
    (i) Viruses are non-cellular and ultramicroscopic.
    (ii) Virus has two components
        (a) A core of nucleic acid called nucleoid.
        (b) A protein coat called capsid.
           Envelope, only in                        Capsomeres, together form
          some larger viruses.                      capsid, a protein coat
                                                    usually highly symmetrical.
                      Genetic material,    Core region
                       DNA or RNA         inside capsid
                         Structure of a virus (generalised)
2. Viroids (RNA without a Capsid)
TO Diener (1917) introduced the term as ‘Subviral pathogens’. Viroids
are 100 times smaller than smallest virus. They are known to be
infectious for plants only (no animal), e.g., potato spindle tuber caused
by viroids.
Virion
An intact, inert, complete virus particle capable of infecting the host
lying outside the host cell in cell free environment is called virion.
Virusoids
These are like viroids, but located inside the protein coat of a true
virus. Virusoid RNA can be circular or linear. These are non-infectious
as they are replicated only in their host.
Prions/Slow Virus
The prions are smallest, proteinaceous infectious particles, i.e., disease
causing agents that can be transmitted from one animal to another.
                                                                                          3
                 Plant Kingdom
Plants : Producers of the Ecosystem
Plants are multicellular, photoautotrophic and embryo forming
(excluding algae) organisms placed in kingdom–Plantae. They have
cell wall, which is made up of cellulose and reserve food material in the
form of starch (sometimes fat as in seeds).
Plants are referred to as producers, because they have unique ability to
fix solar energy in the form of chemical energy, through the process of
photosynthesis. They supply the energy in ecosystem to other living
organisms, hence they are referred to as producers.
The plant kingdom is classified as
                                             Plant Kingdom
       Cryptogamae (non-flowering)                                    Phanerogamae (flowering)
               Thallophyta
                    (plant body is not divided                   Gymnosperms           Angiosperms
                    into root, stem and leaves)               (naked seeded plants)   (covered seed)
       Algae          Bryophyta              Pteridophyta
(non-embryophytes,    (these are          (these are embryo         Monocotyledons Dicotyledons
lack seeds and      embryophytes         bearing plants which
vascular tissue.)  without vascular     form seed and contain
                       tissues.)        vascular tissue as well.)
                                                  Ferns
           Liverworts Hornworts Mosses
Algae (L. Alga–sea weeds)
These are eukaryotic, autotrophic (holophytic), chlorophyll containing,
non-vascular thallophytes. These are characterised by the absence of
embryonic stage and presence of non-jacketed gametangia. Mostly,
they are of aquatic habitat (both freshwater and marine).
The branch of Botany which deals with the study of algae is termed as
‘Algology or Phycology’. FE Fritsch is known as ‘Father of Algology’.
(Prof. MOP Iyengar is regarded as Father of Indian Algology).
                                                           Plant Kingdom         25
Classification of Algae (FE Fritsch; 1935)
   Algal Class          Colour          Reserve Food              Examples
Chlorophyceae       Grass green        Starch             Chlamydomonas and
                                                          Spirogyra.
Xanthophyceae       Yellow-green       Fat                Microspora and Botrydium.
Chrysophyceae       Yellow-green and   Carbohydrate and   Amphipleura and
                    golden-brown       leucosin           Chrysosphaera.
Bacillariophyceae   Brown and green    Fat and volutin    Pinnularia and Melosira.
Cryptophyceae       Red and            Carbohydrate and   Cryptomonas.
                    green-blue         starch
Dinophyceae         Dark yellow,       Starch and oil     Peridinium and
                    brown-red                             Glenodinium.
Chloromonadineae Bright green          Fatty compounds    Vaucheria and Trentonia.
Euglenophyceae      Grass green        Paramylum          Euglena and Phacus.
Phaeophyceae        Brown coloured     Laminarin and      Laminaria and Fucus.
                                       mannitol
Rhodophyceae        Red coloured       Floridean starch   Polysiphonia and
                                                          Batrachospermum.
Myxophyceae         Blue-green         Protein granules   Nostoc and Anabaena.
Characteristics of Algae
Important characteristics of algae are given below
Structure
Algae may be unicellular and multicellular.
1. Unicellular
It is of two types
    (i) Motile, e.g., Chlamydomonas, etc.
   (ii) Non-motile, e.g., Chlorella, etc.
2. Multicellular
It is of following types
     (i) Colonial, e.g., Volvox, Hydrodictyon, etc.
    (ii) Aggregation, e.g., Tetraspora, Prasinocladus, etc.
   (iii) Filamentous, e.g., Ulothrix, Cladophora, etc.
   (iv) Pseudoparenchymatous, e.g., Nemalion, etc.
    (v) Siphonous, e.g., Vaucheria, etc.
   (vi) Parenchymatous, e.g., Ulva, Fritschiella, etc.
  (vii) Well-developed thallus, e.g., Chara, Sargassum, etc.