Biology 11 Reviewer
2nd Long Exam
Roots
   Germination
   o Epigeal  cotyledons and shoot apex emerge
      while hypocotyl elongates
   o Hypogeal  cotyledons remain within testa
      and hypocotyl elongates only slightly or not at
      all
   Types of root system
   o Taproot system
   o Fibrous root system
   o Adventitious roots (cassava, sweet potato)
                                                        pericycle  origin of phloem and xylem
                                                        xylem differentiation in primary roots
                                                        o from periphery inwards (centripetal)
                                                        o exarch
                                                        protoxylem first before metaxylem
                                                        Casparian strips
                                                        o in endodermis
                                                        o impermeable to small molecules to prevent
                                                            diffusion of ions from the apoplast within the
                                                            vascular column
                                                        Lateral root primordial
                                                        o continue to develop and grow out through the
                                                            cortex and ruptures through the epidermis
                                                        o becomes a fully developed root by the time it
                                                            emerges
   As cells enlarge, they begin to show
   1. differences in wall thickening
   2. cell shape, cell contents
   3. differences in activity and function
   Rates of pattern formation
   o rapidly-growing seedlings (corn or pea)  fully
       matured primary tissues 10-15 mm behind the      Secondary growth
       tip.                                             o occurs when cells in the residual procambium
   o slow-growing roots  mature tissues may be            and later parts of the pericycle begin to make
       found within a few mm of the apex                   periclinal divisions.
   o   Only the pericyle cells opposite the xylem
       points start to make periclinal divisions.
   o   inner layer: vascular cambium.
   o   outer layer: pericycle.
    Formation of vascular cambium
    o Layers (from outermost):
             Cortex
             secondary phloem
             vascular cambium
             secondary xylem
    Formation of periderm
    o periderm replaces the epidermis.
    o develops from the activity of the phellogen or
        cork cambium
NOTE: cork cambium originates from the division of
the pericyle
Stems                                                    Primary tissues in the stem
   Leaves can drop off, leaving a scar but there will    o Origin: protoderm, procambium and ground
   still be a bud or shoot into which it has developed       meristem
   As shoot apex grows, older leaf primordia are left           Epidermis; ordinary epidermal cells, guard
   behind.                                                      cells, idioblasts, trichomes
   New primordia are continuously borne above                   Cortex: parenchyma, collenchyma may be
   existing ones                                                continuous or separated
                                                                     strips). In others may contain
                                                                sclereids, secretory cells, laticifers.
                                                                Endodermis: conspicuous in stems of
                                                                lower vascular plants`
                                                         Primary vascular system
                                                         o   In gymnosperms and dicotyledons it is a
                                                             continuous or split cylinder. If split, each
    Specialized stems                                        strand is called vascular bundle.
    o Rhizomes  horizontally growing                             Collateral bundle: phloem is external to
    o Bulbs  vertically-growing underground roots                xylem
        with fleshy leaves and short stem
                                                                  Bicollateral bundle: phloem on the inner
    o Stolons  horizontally growing with plantlets
        along nodes                                               side of xylem.
    o Tubers  enlarged tips of rhizomes                 Vascular cambium - a cylinder of meristematic
    o Runners                                            cells that forms secondary vascular tissue.
            in grasses; horizontal                       o The accumulation of this tissue increases the
            when these branches touch the soil they          diameter of a woody plant.
            produce adventitious roots.
                                                         o Secondary xylem - forms to the interior
    o Taro (gabi) corm
            vertically growing enlarged stem             o Secondary phloem  forms to the exterior of
    Primary growth                                           the VC
    o Apical meristem -> Primary meristem
            Protoderm -> epidermis
            Procambium -> stele, vascular cambium
            Ground -> cortex
                                                         NOTE: trees do not need a heartwood to survive
                                                         In temperate countries:
                                                         o Activity of the cambium is commonly
                                                              periodic. The xylem produced during one
    growth period constitutes a growth layer
    (growth or annual rings).
Parts of the periderm
o Phellogen
         cork cambium, a lateral meristem
         arise from epidermis, cortex or phloem
o phellem
         in compact rows with no intercellular
         spaces
         Primary wall with cellulose, sometimes
         with lignin or suberin
         Suberin layer is impermeable to water and
         gases and can resist action of acids.
         Are dead cells
o Phelloderm
         living cells with non-suberized walls
         may store starch, in others contain
         chloroplasts and photosynthetic
         Sclereids may be present.
first cork cambium  may form as epidermal cells
o Inner cells  cork cambium
o Outer cells  cork
Lenticel- protrude above surrounding periderm
because of bigger size and loose arrangement of
the cells
Leaves                                                   o Betalains  contain nitrogen
                                                         o Anthocyanins  no nitrogen
    Primary growth of shoots                             Tendrils (cadena de amor, grape, pisum)
    o As shoot apex grows, older leaf primordia are      o is a specialized stem, leaf or petiole with a
        left behind.                                         threadlike shape that is used by climbing
    o New primordia are continuously borne above             plants for support and attachment, generally by
        existing ones.                                       twining around whatever it touches.
    o In angiosperms specially, as young                 o Can photosynthesize
        primordium develops from SAM, preparations       o Can be formed from modified shoots,
        are under way for emergence of the next              modified leaves
        primordium                                       Nepenthes
    Types of leaves                                      o An epiascidiate leaf
    o Palmately compound leaf                            o If meristematic activity continues the lamina
    o Pinnately compound leaf (rachis, leaflets)             can become funnel-shaped forming a container
    o Simple leaf - blade consists of only one piece     FROM EXERCISE 7:
    o Parallel monocotyledon leaf (blade, auricle,       o Phyllotaxy  system of leaf arrangement on
        sheath)                                              the stem
    Gymnosperm (pine leaf)                                        Alternate or spiral  one leaf per node
    o Epidermis- heavily cuticularized & with thick               Opposite  two leaves per node
        walls                                                     Whorled  three or more leaves per node
    o Hypodermis (beneath epidermis)  fiberlike         o Blade/Lamina  thin, flattened, green structure
        cells with thick walls.                          o Petiole  holds the blade upright
    o Lobed mesophyll cells                                       Petiolules  stalk of each leaflet
    o Resin ducts                                        o Stipules  outgrowths in pairs found at the
    o Sunken stomata                                         base of some dicot leaves
    o Transfusion tissue                                          Stipulate and exstipulate
    Dicots                                                        Stipels  outgrowths in pairs at the base of
    o Netted venation                                             the petiolule
    o palisade layer occurs adaxially (upper),           Venation
        spongy layer occurs abaxially (underside)        o Netted  veins branch profusely and form a
NOTE: Isobilateral leaf- A leaf in which the palisade        network over the blade (dicot except plantain)
                                                         o Parallel  veins do not form a network
parenchyma occurs adaxially and abaxially beneath
the epidermis of the leaf                                    (monocot except gabi and ube)
                                                         Internal structure of leaves
    Monocot                                              o Epidermis  outermost layer, with cuticle, with
          o     Bundle sheath or border parenchyma           stomata and guard cells, more stomata on
          o     Bulliform cells  thin-walled, highly        lower epidermis than upper epidermis
                vacuolated water-containing cells        o Mesophyll  photosynthetic parenchymatous
            Rapid expansion of leaf: expansion tissues       region between upper and lower epidermis
            Changes in turgor in opening/closing of      o Palisade layer  elongated and closely
            leaves: motor cells                              arranged (perpendicular to upper epidermis)
    Modified leaves                                      o Spongy layer  loosely arranged and
    o Spines                                                 irregularly-shaped (for gas exchange)
    o Storage leaves                                     o Vascular tissues  form vascular bundles in
    o Reproductive leaves                                    leaves
    o Bracts                                                     Border parenchyma  thin-walled cells
            Bougainvillea  colored bracts (get their            Bundle sheath  thick-walled cells
            color from betalains)                        o Bulliform cells  large thin-walled cells in
    Flowers                                                  monocot for rolling in of leaves
    o Pigments present are called betalains
    o Betalains and anthocyanins are water-soluble
       pigments found in the vacuoles of plant cells
                                                   Border parenchyma (thin-walled cells in dicots)
                                                   Bundle sheath (thick-walled cells)
                                                   Transpiration
                                                   o Evaporation of water from from aerial parts of
                                                      the plant
                                                           Stomatal transpiration
                                                           Cuticular transpiration
                                                           Lenticular transpiration
                                                   o Factors
                                                           Light intensity*
                                                           Wind velocity *
                                                           Temperature
                                                           Humidity
                                                           Soil Water Supply
                                                   Guttation
                                                   o Secretion of water to the surface of leaves
                                                      through specialized pores called
                                                      HYDATHODES
Pine needle
After boiling, only sclerenchyma and vascular
tissues remain (petiole, leaf veins, and midrib)
Leaf
o Epidermis
         Cuticle
         Stomata and guard cells
         Bulliform cells in monocots
o Mesophyll
         Palisade layer
         Spongy layer
         Vascular tissues