"Dryosauridae is a monophyletic clade of small, cursorial, plant-eating ornithopod dinosaurs that lived in both Gondwana and Laurasia. This wide distribution is particularly evident during the Late Jurassic with the Laurasian Dryosaurus and Eousdryosaurus, and the Gondwanan Dysalotosaurus. The dryosaurid record from the Upper Jurassic of Portugal is based mainly on isolated material from the Lusitanian Basin in the central-west of the country. So far, merely a partial specimen from the Porto das Barcas locality represents Eousdryosaurus nanohallucis. In this context, the discovery of new localities bearing new dryosaurid specimens is worthy of note. Two new localities have provided only isolated femora, whereas a third comprises a well-preserved partial skeleton of a small dryosaurid ornithopod. The former specimens come from the upper Kimmeridgian-lower Tithonian Praia da Amoreira-Porto Novo Formation and the Tithonian Freixial Formation. The most complete specimen is a partial skeleton from the Tithonian beds of the Bombarral Formation in Peniche. It comprises mainly appendicular bones (humerus, femur, tibia) of a single individual. All the femora recorded exhibit two dryosaurid synapomorphies such as the proximally placed fourth trochanter, and the scar for the M. caudi femoralis longus restricted to the medial surface of the femoral shaft insertion and widely separated from the fourth trochanter. The new evidence of these dryosaurid ornithopods indicates that these small cursorial dinosaurs were common inhabitants and played an important role within the Iberian herbivorous communities during the Late Jurassic."
Abstract: The diplodocid record from the Upper Jurassic of the Lusitanian Basin is relatively scarce. This record includes Dinheirosaurus and some fragmentary remains, among which there has been proposed a second taxon. The study of three new specimens from Valmitão (Lourinhã), Cambelas (Torres Vedras) and Praia Vermelha (Peniche) and the redescription of Dinheirosaurus is providing new information about the diplodocids of the Lusitanian Basin. Preliminary phylogenetic analyses suggest that these new specimens and Dinheirosaurus are derived diplodocids closely related to Diplodocus and Barosaurus from the Morrison Formation.
Resumo: O registo fóssil de diplodocídeos proveniente do Jurássico Superior da Bacia Lusitânica é relativamente escasso. Este registo inclui Dinheirosaurus e outros exemplares mais fragmentários, dos quais se considera a existência de um segundo taxon. O estudo de três novas ocorrências em Valmitão (Lourinhã), Cambelas (Torres Vedras) e Praia Vermelha (Peniche) assim como a redescripção de Dinheirosaurus têm fornecido nova informação sobre os diplodocídeos da Bacia Lusitânica. Análises filogenéticas preliminares sugerem que estes três espécimenes e Dinheirosaurus correspondem a diplodocídeos derivados relacionados com Diplodocus e Barosaurus da Formação de Morrison.
GEOGRAPHIC AND STRATIGRAPHIC DISTRIBUTION OF THE SAUROPOD TURIASAURUS AND TURIASAURIA CLADE
Rafael Royo-Torres, Alberto Cobos, Francisco Gascó, Pedro Mocho, Francisco Ortega, Luis Alcalá.
"Turiasauria clade represents a distinct group of non-neosauropods with a wide geographic distribution across Europe and probably Africa during the Middle-Late Jurassic. This clade is known thanks to the study of the sauropod dinosaur Turiasaurus riodevensis Royo-Torres, Cobos & Alcalá, 2006. It was discovered in 2003 in Riodeva, (Teruel province, South of Iberian Range, Spain). Turiasaurus is defined by the representative cranial and postcranial remains found in its type-locality: skull, mandible, eight teeth, six cervical vertebrae with ribs, four dorsal vertebrae, eight dorsal ribs, the sacrum, two distal caudal vertebrae, a proximal fragment of the left scapula, a left sternal, a complete left forelimb, fragments of ilium and ischium, a left pubis, a distal fragment of the left femur, a proximal and a distal fragments of the left tibia, a left fibula, two astragali and a pes. In addition, Riodeva area has yielded 3 other specimens of turiasaur sauropods. From one of them, the San Lorenzo specimen, there are cranial and postcranial remains, which are currently being studied. All the sites with turiasaur remains are placed in the Villar del Arzobispo Formation (dated Upper Kimmeridgian-Berriasian). Aside from the specimens of Riodeva, we tentatively assign materials from other localities to Turiasauria clade. Phylogenetic analyses support the attribution to Turiasauria of three sauropods: one from the Middle Jurassic of Beni Mellal (Moroco), Atlasaurus imelakei, and two species from the Upper Jurassic of Spain, Galveosaurus herreroi from Galve (Teruel) and Losillasaurus giganteus from Losilla (Valencia). Besides these taxa, and according to some synapomorphies, different specimens have been considered like potential turiasaurs. Turiasaurus possess characteristic ‘heart’-shaped teeth in labial profile with their apex labiolingually compressed, and with an asymmetrical shape produced by a concave distal margin near the apex even when unworn. This type of teeth allows the inclusion in Turiasauria of the Middle Jurassic of England tooth referred to Cardiodon, the Middle Jurassic of Peterborough (England) teeth assigned to “Cetiosauriscus leedsi”, four teeth from the Upper Jurassic of Aylesbury (England) assigned previously to “Hoplosaurus” and “Pelorosaurus”, some teeth from the Upper Jurassic of Asturias (Spain), and the teeth assigned to “Neosodon” in France. In Portugal, more than 26 Oxfordian-Tithonian teeth have been included in Turiasauria. Some of them are associated to an articulated forelimb and chevrons from the Lourinha Formation of Portugal. In Africa in addition to Atlasaurus, some of the postcranial material from Tendaguru beds (Kimmeridgian-Tithonian) in Tanzania has also been referred to Turiasauria, i.e. the caudal series of the specimen MB.R. 2091.1-30 and the right manus MB.R. 2093.1-12 (Museum für Naturkunde in Berlin). Thus, at present, Turiasauria is proposed as a distinct clade of sauropods that diverged, prior to the Middle Jurassic, from the lineage leading to neosauropods and which spread across Europe and Africa during the Late Jurassic. Turiasaurus represents a very large sauropod (over 25 m in length), suggesting that gigantic body sizes were achieved not only by neosauropod clades, such as Diplodocidae and Titanosauriformes, but also by non-neosauropods."------