Biodiversity
-‐
notes:
1. Four
metrics/indices
of
interest:
a. Relative
Abundance
(pi)
i. What
portion
of
the
total
population
(N)
is
one
particular
species
(ni)
ii. pi
=
ni
/
N
iii. For
example,
two
communities
may
both
contain
the
same
number
of
species
but
one
community
may
be
dominated
by
one
species
while
the
other
community
may
contain
large
numbers
of
all
species
b. Species
Richness
(S)
i. Total
number
of
species
present
within
an
ecosystem
c. Species
Evenness
(J’)
i. how
close
in
numbers
each
species
in
an
environment
are
ii. can
be
represented
by
Pielou's
Evenness
Index:
1. J’
=
H’/ln(S)
a. Where
H’
is
the
Shannon
Diversity
Index
and
S
is
the
total
number
of
species
2. J'
is
constrained
between
0
and
1.
The
less
variation
in
communities
between
the
species,
the
higher
J'
is
d. Species
Diversity
i. A
measure
of
the
diversity
within
an
ecological
community
that
incorporates
both
species
richness
and
species
evenness
ii. Shannon
Diversity
Index
(H’)
1. Used
to
quantify
the
entropy
(uncertainty)
in
population.
The
idea
is
that
the
more
different
species
there
are,
and
the
more
equal
their
proportional
abundances
in
the
ecosystem,
the
more
difficult
it
is
to
correctly
predict
which
species
will
be
the
next
one
selected
at
random.
The
Shannon
entropy
quantifies
the
uncertainty
(entropy
or
degree
of
surprise)
associated
with
this
prediction.
2. Biodiversity
Trends:
a. Latitudinal
Diversity
Gradient
(LDG)
i. In
general,
as
latitude
increases,
biodiversity
decreases
ii. There
exists
a
general
lack
of
consensus
as
to
why
this
pattern
exists
(see
Wikipedia
for
a
good
summary
of
the
major
theories
to
date)
b. Elevational
Diversity
Gradient
(EDG)
i. Species
richness
(S)
tends
to
increase
as
elevation
increases,
up
to
a
certain
point,
creating
a
"diversity
bulge"
at
middle
elevations
(approx.
1,000m-‐1,300m
[3,200-‐4,300
ft])
c. Rapoport’s
Rule
(Eduardo
Rapoport)
i. States
that
latitudinal
ranges
of
plants
and
animals
are
smaller
at
low
latitudes
and
altitudes
1. Niche
specialization
d. Biodiversity
trends
through
history:
i. See
Phanerozoic
biodiversity
graph
on
course
website
1. Growth
follows
hyperbolic
model,
in
which
second-‐order
positive
feedback
of
an
increase
in
biodiversity
causes
an
increased
growth
in
biodiversity
(goes
beyond
first-‐order
positive
feedback
system
where
resource
availability
alone
allows
for
more
biodiversity)
2. In
the
midst
of
the
Holocene
extinction
event
since
the
emergence
of
humans,
some
arguing
that
current
rate
is
enough
to
eliminate
most
species
on
earth
within
next
100
years