Lesson 07
Lesson 07
                                                                  7
               Notes
                                                   SHOOT SYSTEM
                         Shoot system is an aerial and erect part of plant body which grows upwards. It is
                         usually above the soil and develops from plumule of the embryo. It consists of stem,
                         branches, leaves, flowers, fruits and seeds. In this lesson you will study about the
                         structure, types, modifications and functions of stem, leaf, flower and fruit.
OBJECTIVES
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 7.1 STEM                                                                                                                            Plants and animals
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  Plants and animals     7.1.3 The Shoot Apex
                         Shoot apex is the terminal, dome shaped part of shoot, formed of meristem called
                         apical shoot meristem responsible for the development and differentiation of
                         primary permanent tissue and mainly causes growth in length. It is divided into two
                         regions - Tunica and Corpus (Fig. 7.1)
                                                                                      Tunica
               Notes
                                                                                     Corpus
                                                          (a)                                                 (b)
                                                 Fig.7.1 a-b L.S. of shoot apex to show tunica and corpus
                         (i) Tunica (covering)- An outer zone of shoot apex, 1-3 layers in thickness. It gives
                             rise to epidermis and is responsible for surface growth, and its cells divide only
                             anticlinally.
                         (ii) Corpus (body)- Inner multi-layered zone of cells which divide in all directions.
                              They finally give rise to procambium (forms vascular tissue) and ground
                              meristem (forms ground tissue). These cells also form leaf primordia (a newly
                              developing leaf).
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7.1.5 Types of stem                                                                      Plants and animals
The stem may be (i) aerial (erect, rigid, strong and upright as in herbs, shrubs and
trees) (ii) sub aerial (weak, unable to stay upright and trail on ground as creepers
or climb up as climbers) or (iii) underground (buried in soil and produces aerial
branches under favourable conditions only).
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  Plants and animals     3. Bulb                 Reduced, flattened discoid stem                Onion
                            (Fig.7.2ci,ii)       with crowded nodes bearing
                                                 overlapping fleshy (inner) and
                                                 dry (outer) scale leaves. Terminal
                                                 bud (in centre) forms foliage
                                                 (green) leaves. Adventitious
                                                 roots grow from discoid base.
               Notes
                         4. Tuber                Swollen tips of underground                    Potato
                            (Fig.7.2d)           lateral branches of stem, store
                                                 food as starch, bear “eyes”. Each
                                                 eye is a node which bears bud
                                                 and scar of scale leaves.
                                                               Scale Base
                           Scale leaf        Adventitious        Scale leaf Base of Scale Fleshy
                         Nodes                  Root           Adventitious               Scale Leaf
                                                                                             Apex
                                                                  Root                          Adventitious         Eye
                                                                    Bulb                          Root
Corm
                         Fig.7.2 Underground modifications of stem – (a) Rhizome of Ginger, (b) Corm of Yam, (ci)
                                             Bulb of Onion (cii) V.S. bulb, (d) Tuber of potato.
                         Sub aerial modifications Of stem- Stems are weak, therefore lie prostrate on the
                         ground or may get partially buried in the top soil. The plants bearing such stems
                         are called creepers. Their stems serve the function of vegetative propagation.
                                             Table 7.3 Modifications of Sub aerial stems
                            Type                            Characters                                   Examples
                         1. Runner           Long, weak, slender branch with long internodes.        Grass, Oxalis
                            (Fig.7.3a)       Runs horizontally on soil surface giving off
                                             adventitious roots at nodes
                         2. Stolon           Weak lateral branch which grows upwards then            Mint (‘Pudina’),
                            (Fig. 7.3b)      arches down to meet the soil, strike roots and          Jasmine
                                             produce daughter plants.
                         3. Offset           Like runner but thicker and shorter, grow for a         Water hyacinth,
                            (Fig.7.3c)       short distance then produce cluster (rosette) of        water lettuce
                                             leaves above and adventitious roots below;
                                             generally in aquatic plants
                         4. Sucker           Underground runner which grows horizontally             Chrysanthemum
                            (Fig.7.3d)       for a distance under soil then emerges obliquely
                                             upwards, strikes roots and forms daughter plants
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Aerial stem modifications - Whole stem or its part (axillary or terminal bud) gets
modified to perform definite functions. You can recognise them as stems by
following features :
(i) Arise in the axil of leaf (ii) Bear nodes and internodes (iii) may bear leaves, buds,
flowers.
 1. Stem tendrils        Thread like, spirally coiled, leafless structures   Grape vine
    (Fig.7.4a)           (tendrils) which twine around neighbouring
                         objects and help weak plants to climb
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  Plants and animals          Apical tendril
                                                                                                       Leaf spine
                                                                                                         Phylloclade
                                                                                                              Flower                 Cladode
                                                               Thron
               Notes
                         Apical
                         tendril                                                                                                                  Hook
                         pushed to
                         side
                         1. What are plants with weak stem trailing on the ground known as?
                             ............................................................................................................................
                         2. Name the modification to which Runner, Stolon, Offset and Sucker belong.
                             ............................................................................................................................
                         3. What is a phylloclade with one or two internodes called?
                             ............................................................................................................................
                         4. ‘Haldi’ and onion belong to which category of stem modification respectively?
                             ............................................................................................................................
                         5. Match the items of column A with those of column B
                                            A                                               B
                             (a) Tendril                                         (i) Protection
                             (b) Sucker                                         (ii) Perennation
                             (c) Thorns                                        (iii) Reproduction
                             (d) Bulb                                          (iv) Photosynthesis
                             (e) Phylloclade                                    (v) Climbing
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2. Conduct water and minerals from roots to leaves and manufactured food from                                                       Plants and animals
   leaves to different parts of the plant.
3. Bear flowers and fruits
B. Secondary Functions
1. Storage - Stems store food and water in some plants e.g. potato
2. Perennation - The underground stems help tide over the unfavourable growing Notes
   periods e.g. ginger.
3. Vegetative propagation - Stem can be a means of vegetative propagation e.g.
   rose, and sugarcane.
4. Photosynthesis- in certain plants like xerophytes (desert plants) where leaves
   are reduced, the stem takes up the function of photosynthesis. These stems
   possess chlorophyll e.g. Opuntia
5. Protection- In some plants the axillary bud modifies into thorn and protects the
   plants from grazing animals e.g. citrus, Duranta.
6. Climbing - Tendrils or hooks are modified branches or buds. They coil around
   the support and help the plant to climb e.g. grape vine
                                                                                                  Epidermis
                                                                                                  Hypodermis
                                                                                                  (Collenchyma)
                                                                                                                     Pericycle Cortex
                                                                                                 General Cortex
                                                                                                 (parenchyma)
                                                                                                 Endodermis
                                                                                                 Sclerenchyma
                                                                                                 (Hard Bast)
                                                                                                 Phloem
                                                                                                                  Vascular Bundle
                                                                                                 Cambium
                                                                                                 Metaxylem
                                                Epidermis                                        Protoxylem
                                                Cortex
                                                Primary phloem
                                                                                                 Wood
                                                                                                 Parenchyma
                                                 Primary xylem
                                                                                                 Pith
                                  (a)               Pith
                                                                                   (b)
                                        Fig. 7.5 T.S. Dicot stem. a-Diagrammatic b-A portion enlarged.
                            (iii) Medullary rays - Narrow regions of parenchymatous cells in between the
                                  vascular bundles.
                            (iv) Pith - The central parenchymatous zone with intercellular spaces.
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B. Internal structure of monocot stem (e.g., maize)                                         Plants and animals
A transverse section of monocot stem reveals the following structures (7.6a and b)
1. Epidermis - Single layered, covered with cuticle, stem hairs absent.
2. Ground tissue- A mass of parenchymatous tissue. Only a few peripheral layers
   below epidermis are sclerenchymatous called hypodermis.
     1. Vascular bundle- Numerous, scattered in the ground tissue each enclosed Notes
        by sclerenchymatous bundle sheath. Each bundle is
        (a) collateral and (b) closed (no cambium strip between xylem and
        phloem) with (c) endarch xylem. Xylem occurs in the form of letter ‘Y
        ‘and innermost protoxylem disintegrates to form a water cavity.
                          Epidermis
                          Hypodermis
                            (Sclerenchyma)
                             Vascular bundles
                                                        Crushed Phloem
Ground tissue
          Fig. 7.6 T.S. Monocot stem. (a) Diagrammatic (b) A portion enlarged
                           (c) A vascular bundle magnified.
Anatomical differences between dicot and monocot stem, and anatomical differences
between root and stem are given in Tables 7.5 and 7.6
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  Plants and animals                 Table 7.5 Differences between monocot stem and dicot stem
                                 Characters                               Dicot stem                                      Monocot stem
                          1. Epidermal hairs                    Present                                        Absent
                          2. Hypodermis                         Collenchymatous                                Sclerenchymatous
                          3. Ground tissue                      Differentiated into cortex,                    Undifferentiated
                                                                endodermis, pericycle, pith
               Notes                                            and medullary rays
                          4. Vascular bundles                   (i)     Number not very large                  (i)      Numerous
                                                                (ii)    Uniform in size                        (ii)     smaller near periphery,
                                                                                                                        bigger in the centre
                                                                (iii) arranged in a ring                       (iii) scattered
                                                                (iv) open                                      (iv)     closed
                                                                (v)     bundle sheath absent                   (v)      bundle sheath present
                                                                (vi) xylem vessels arranged                    (vi)     xylem vessels arranged
                                                                     in a radial row                                    in shape of letter “Y”
                                                                (vii) water cavity absent                      (vii) water cavity present
                          5. Secondary growth                           Present                                         Mostly absent
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4. If you want to study the internal structure of a monocot and a dicot stem, name                                                   Plants and animals
   the plants you would select for the study.
    ............................................................................................................................
7.1.9 Secondary growth in stem
You have learnt in lesson 6 about the secondary growth in dicot roots and its
importance, let us study it in stem. It occurs only in dicot stem a little away from
the shoot apex and helps the plant to (a) grow in girth (thickness) and (b) makes Notes
it very strong to stand upright for many years. That is why you see that very tall
trees can withstand strong winds, and lashing rains without falling down but
monocot plants like wheat, rice, maize, and grasses bend easily due to absence of
secondary growth in their stems.
Growth in thickness in dicot stem becomes possible due to the formation of new
tissues entirely by the activity of two lateral meristems -(i) Vascular cambium
and (ii) Cork cambium (Fig.7.7 a-d).These tissues thus formed are known as
secondary tissues and growth in girth is referred as secondary growth.
(i) Activity of vascular cambium -Forms secondary vascular tissues as follows
         z The strip of cambium present in the vascular bundle is called Fascicular
           Cambium (Fig 7.7a)
         z The cells of medullary rays adjoining the strip of vascular (Fascicular) cambium
           become meristematic and form interfascicular cambium( Fig. 7.7b).
         z Both fascicular and inter-fascicular cambium join to form a continuous
           cambium ring (Fig. 7.7b,c)
                                                                                   Med rays              Phellem or cork
                 Epidermis                                                        Secondary                    Phellogen
                   Cortex                                                           xylem
              Primary Phloem
             Fascicular cambium
                Primary-xylem
                                                                       Secondary phloem
                      Pith         Interfascicular cambium                Cambium ring
                     (a)                    (b)                    (c)             (d)
      Fig. 7.7 (a-d) T. S. Dicot Stem- Various stages in secondary growth (Diagrammatic)
         zCambium divides and adds cells on internal side (towards pith) which
          mature into secondary xylem and cells added towards external side
          (periphery ) mature into secondary phloem (Fig 7.7c).
      z Amount of secondary xylem produced is comparatively comparatively
          more than secondary phloem (Fig7.7d)
(ii) Activity of cork cambium-Forms periderm as follows :
      z Cork cambium or phellogen develops in the cortex.
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  Plants and animals           z   Phellogen divides and adds cells on both the inner and the outer side.
                               z   Cork cells are compactly arranged and become dead and suberized
                                   (deposition of suberin) except in regions of lenticels (Fig. 7.8) where cells
               Notes               are loosely arranged (complimentary cells) and non-suberized. It is
                                   through the lenticels that woody branches and tree trunks can undergo
                                   gaseous exchange.
                               z   All the dead cells lying outside the active phellogen constitute the bark.
                                                                           Lenticel
                                                                                    Bark
                                                                           Epidermis
                                                                           Cork
                                                                           Cork-cambium Periderm
                                                                           Secondary Cortex
                                                                           Collenchyma
                                                                           Cortex
                                                                           Sclerenchyma
                                                                           Secondary
                                                                           Phloem
                                                                           Cambium
                                                                          Autumn wood
                                                                          Secondary
                                                                          Medulary Second
                                                                          Ray       Annual
                                                                                     Ring
                                                                                          Secondary
                                                                         Spring wood        Wood
                                                                         Autumn Wood
                                                                                      First
                                                                        Secondary
                                                                                      Annual
                                                                        medullary ray
                                                                                      Ring
                                                                        Spring wood
                                                                        Primary wood
                                                                        Pith
                                               Fig. 7.8 T.S. of old stem, A Portion enlarged
                         In Betula bhojpatra bark peels off like sheets of paper. Ancient manuscripts are still
                         preserved on them. Cork tissue becomes very thick in Cork tree (Quercus suber)
                         and is used commercially as, bottle-stoppers, insulators, and shoe soles.
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    ............................................................................................................................
3. Define bark.
    ............................................................................................................................
4. Why are lenticels, non suberized?
    ............................................................................................................................
5. The stems of grasses, and rice, remain weak and thin, why?
    ............................................................................................................................
6. Which layers constitute the periderm? What is it’s function?
    ............................................................................................................................
7.1.10 Wood
Wood is the secondary xylem produced by the activity of vascular cambium in dicot
stem.
Annual Rings (A secret to know the age of tree)
In temperate regions, the climatic conditions show pronounced seasonal variations.
The activity of vascular cambium also becomes periodical as a result, distinct growth
layers are formed in xylem. In spring season cambium is very active and produces
a greater number of vessels with wider cavities. The wood formed during spring
is called early wood (or spring wood). In summer, cambium is less active and forms
narrow vessels, this wood is called late wood (or summer wood). These two kinds
of woods in a transverse view appear as alternate concentric rings together forming
an annual ring (Fig 7.8). By counting the number of these annual growth rings
we can know the age of a tree. Science dealing with predicting the age of a tree
by counting the annual growth rings is called as Dendrochronology.
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  Plants and animals                                                                          Phloem
Bark Sapwood
Heartwood
               Notes
                                                                                             Annual rings
Xylem rays
                                                Fig. 7.9 T.S. old stem showing Heart wood and Sap wood.
                         Table 7.7 enlists the main differences between sap wood and heart wood.
                                   Table 7.7 Differences between sap-wood and heart wood
                                          SAP WOOD                                                HEART WOOD
                          1. It is the outer light coloured wood of                        It is the central dark colured wood of an
                             an old stem                                                   old stem
                          2. Light coloured                                                Dark coloured due to presence of gums,
                                                                                           resins, oils, tannin etc.
                          3. Contains living cells                                         Living cells are absent
                          4. Vessels not plugged and help in upward                        Vessels are plugged with tyloses
                             movement of water and minerals
                          5. Wood is lighter in weight                                     Heavier in weight
                          6. Less durable because of susceptibility                        More durable, resistant to attack of the
                             to attack of pathogens                                        pathogens
                          7. Commercially less valuable                                    Commercially more valuable
                         Mechanical tissues in stem -The stem of a tall tree needs to i) resist against pulling
                         forces of wind and ii) to stand erect against gravity. Stem gets this strength from
                         - Sclerenchyma in hypodermis and it’s patches in the pericycle and secondary
                         phloem, abundant lignified vessels, tracheids and fibres in secondary xylem i.e. wood
                         and sclereids in pith.
7.2 LEAF
Leaf is a flattened and expanded lateral appendage of stem or branch developing Notes
from its node. It originates from leaf primordium formed by the shoot meristem and
bears a bud in its axil called axillary bud. It is the seat of very important
physiological processes like photosynthesis, transpiration and respiration. Besides
protecting axillary buds ,leaf can get modified into structures for storing food and
water, climbing, and vegetative propagation.
                          Leaf apex
                        Leaf margin
Mid rib
Lamina
Fig. 7.9 Leaf and its parts Fig. 7.10 Variations in leaf shape.
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Notes
Fig. 7.11 Variations in Leaf apices Fig. 7.12 Variations in Leaf margins
                                         Reticulate
                                         network
                                                            Strong veins
                                                                                       (a)
                             Pinnate                        from pellicle                                    (b)
                               (a)              Palmate
                                                      (b)
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1. Define venation.
    ............................................................................................................................
2. Differentiate between unicostate and multicostate venation.
                                                                                                                                   Notes
    ............................................................................................................................
3. What is the type of venation found in peepal and palm leaves?
    ............................................................................................................................
4. Name the structure which arises in the axil of leaf
    ............................................................................................................................
5. What is the prominent vein called which is present in the middle of lamina and
   runs from base to apex?
    ............................................................................................................................
 1. The leaf has a single undivided                              The lamina is divided into many
    lamina (Fig. 7.9)                                            segments called leaflets (Fig. 7.14)
 2. If divided, the incisions do not touch                        Incisions touch the mid rib (Fig. 7.15)
    the mid rib (Fig. 7.13d)
 Types of Compound leaves - They are of two types as shown in table 7.8
                                 Table 7.8 Types of compound leaf
                       Pinnate                                                   Palmate
 1. Leaflets are attached to mid rib or rachis                   Leaflets radiate from the end of petiole
    and are arranged laterally (Fig 7.15)                        like fingers of a palm (Fig. 7.14 )
 2. Leaflets and mid rib may get further                         Depending upon the number of leaflets
    divided to form compound leaves that                         compound leaves are bifoliate, trifoliate,
    are unipinnate, bipinnate, tripinnate and                    quadrifoliate and multifoliate (Fig.7.14)
    decompound (Fig. 7.15)
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               Notes         Axillary
                                                                                                           Quadrifoliate
                             bud                                                            Trifoliate                          Multifoliate
                                                                          Bifoliate
                                            Palmately
                                         compound leaf
                                                                                                                                     Decompound
                         Axillary bud                           Paripinnate Imparipinnate Bipinnate Tripinnate
                                          Pinnately                   Unipinnate
                                      compound leaf
                                      ACTIVITY 7.1
                         Aim - To collect and study a few leaves.
                         Material required — Collect leaves of peepal, neem, banana, palm, rose, grass,
                         imli and tulsi.
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Method - Observe the following features in the collected material                           Plants and animals
7.2.4 Phyllotaxy
It is the arrangement of leaves on stem or branch. The orientation and arrangement Notes
of leaves is such that they get appropriate amount of sunlight for photosynthesis.
It is of three types
(i) Alternate (Fig. 7.16d) - a single leaf arising at each node e.g. china rose,
    mango.
(ii) Opposite (Fig. 7.16a-b) - Leaves occur in pairs at each node. This arrangement
     may be
    (a) Decussate (Fig. 7.16a) - When the successive pairs of leaves at upper and
        lower nodes are at right angles e. g .,“Tulsi”, Calotropis
    (b) Superposed (Fig. 7.16b) - when the successive leaf pairs at upper and
        lower nodes are exactly in the same plane e.g. guava
(iii) Whorled (Fig. 7.16c) - When there are more than two leaves at each node
      arranged in a circle or whorl e.g. Nerium.
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  Plants and animals                                       Table 7.9 Modifications of leaves
                                     Type                                         Characters                                        Examples
                          l. Leaf Tendril               Here leaves or leaflets get modified to                             Pea, Glory lily
                             (Fig.7.17a)                form thin wiry, closely coiled sensitive
                                                        structure called the tendril that helps the
                                                        plant to climb the support.
               Notes      2. Spines                     The leaves are modified into sharp                                  Prickly poppy
                             (Fig 7.17b)                and pointed structures which protect                                (Argemone)
                                                        the plant and help in reducing                                      Opuntia, Aloe
                                                        transpiration.
Bladder
Pitcher
                                           Fig. 7.17 Modifications of Leaf a-d (a) Leaf tendril; (b) Spines;
                                                         (c) Pitcher plant; (d) Bladderwort
                         Heterophylly (heteros = different)- Some plants show more than one type of leaves
                         in the same plant , this phenomenon is called heterophylly. It is found in some plants
                         which remain partly submerged in water e.g. Water chestnut, and Limnophila
                         1. What is the type of phyllotaxy found in mango, ‘tulsi’ and guava plants?
                             ............................................................................................................................
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2. Match the following items of column A with those of column B                                                                        Plants and animals
                 A                                                         B
     (a) Pitcher                                       (i) Photosynthesis
    (b) Spines                                        (ii) Climbing
     (c) Phyllode                                   (iii) Trapping insects
    (d) Tendril                                      (iv) Protection
3. Give two examples of insectivorous plants.                                                                                    Notes
    ............................................................................................................................
4. Water chestnut shows two different types of leaves on the same plant, what is
   such a condition known as?
      ............................................................................................................................
                                                                                                       Upper epidermis
                                                                                                        Palisade
                                                                                                        parenchyma
                                                                                                     Bundle sheath
                                                                                                     Xylem
                                                                                                       Phloem
                                                                                                          Spongy
                                                                                                          parenchyma
                                                                                                          Sub-stomatal
                                                                                                          cavity
                                                                                           Stoma     Lower epidermis
                                                          Bundle sheath extension
Upper epidermis
Bundle sheath
                         Xylem
                                                                                                         Vascular bundle
                         Phloem
                                                                                                         Mesophyll tissue
                                                                                                          Lower epidermis
                                  Sclerenchyma                 Stoma       Sub-stomatal chamber
                                                 Fig. 7.19 V.S. of a Monocot (Isobilateral) Leaf
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Structure of stomatal apparatus : In dicot leaves, stomatal apparatus is made up                         Plants and animals
of two semi circular guard cells surrounding a pore-stoma (Fig. 7.21) The guard
cells contain chloroplasts and regulate the opening and closing of stomata. Stomatal
pore opens into the inter cellular spaces (substomatal cavity) of mesophyll (Fig.
7.19). The number, shape and distribution of stomata vary (Table 7.10) depending
upon the plant whether it is xerophyte or mesophyte.
                       Table 7.10 Distribution of stomata.
                                                                                                       Notes
     Plants                    Stomatal characters                            Examples
1. Dicots                  Guard cells semicircular reniform             Mango, neem
                           occur generally on lower surface
2. Monocots                Guard cells dumbbell                          Maize
                           shaped, occur on both the surfaces
3. Xerophytes              To reduce transpiration-                      Nerium
                           (i) occur only on lower surface,
                           (ii) are absent or less in number on
                                the upper surface
                           (iii) may be sunken
4. Hydrophytes
   – with floating         Occur only on upper surface                   Lotus
     leaves
   – with submerged        Stomata absent                                Hydrilla
     leaves
Now you can compare the internal structures of dicot and monocot leaves from
Figs. 7.18-19 and Table 7.11
        Table 7.11 Difference between internal structure of Dicot and
                                Monocot Leaf
       Tissue                   Dicot leaf                         Monocot leaf
                             (Dorsiventral leaf)                 (lsobilateral leaf)
1. Epidermis
   (i) Stomata          Occur generally in lower            Occur both in upper and
                        epidermis                           lower epidermis
   (ii) Bulliform cells Absent                              Present in upper epidermus
2. Mesophyll            Differentiated into palisade        Only spongy parenchyma
                        and spongy parenchyma               present
3. Vascular system      (i) in the form of network          (i) in rows
                        (ii) vascular bundle in mid rib     (ii) vascular bundle of midrib is large,
                             region is large, rest of the        but other vascular bundles are
                             vascular bundles decrease           small generally of same size.
                             in size towards the leaf-
                             margin.
B. Special features
(i) Bulliform Cells (Fig 7.19)
      z     These are special type of cells (motor cells) found on upper leaf surface
            of some monocots (e.g. maize, bajra, jowar).
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  Plants and animals           z     They help the leaf to roll and unroll due to change in their turgidity.
                               z     Leaf rolls when these cells lose water due to high rate of transpiration
                                     especially at Mid-day on hot sunny days.
                               z     Thus, under dry conditions they help in reducing the loss of water vapour
                                     through stomata.
                         (ii) Hairs
               Notes
                               z     Hairs are present especially on leaves of plants growing in dry conditions.
                                     They check the rate of transpiration.
                               z     They protect the leaf from bright sunlight, high temperature and air pollution.
                         (iii) Hydathodes (water stomata)
                               z     These are specialised structures (Fig.7.20) present in leaves of angiosperms
                                     (garden nasturtium) occurring in humid climate.
                               z     Through these openings excretion of water and minerals plus simple organic
                                     compounds in liquid form (guttation) takes place. When water absorption
                                     by a plant is more and transpiration is less.
                                                            Guard cell
                                                                Stomatal chamber
                             Inter cellular                                         Guard cells
                                 space                                Epithem                         Epidermal cells
Epidermis
                         Tracheid
                                                                                        Stoma
Chloroplast
Fig. 7.20 Hydathode Fig. 7.21 Stomatal apparatus from a dicot leaf
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 7.3 FLOWER
Flowers are a thing of beauty for us but for the plants they are vital as they are
the seat of sexual reproduction. They produce fruits and seeds.
A flower is a modified shoot because it has (i) nodes very close to one another and
(ii) floral leaves arranged in successive whorls.
7.3.1 Parts of a typical flower (Fig 7.22)
Take a flower of any colour or size growing in your area, you’ll find it’s basic plan
to be the same i.e. the flower is borne on a stalk called pedicel. The pedicel has
a swollen tip known as thalamus or receptacle on which are borne four whorls
successively in definite order as given below :
Accessory whorls
1. Calyx (collection of sepals) - The outer most whorl of green sepals whose main
   function is protection.
2. Corolla (collection of petals) - The next whorl of variously coloured petals. They
   help in attracting insects for pollination.
Reproductive whorls
3. Androecium (male reproductive part) consists of collection of stamens. Each
   stamen has a long slender filament with a bilobed anther at it’s tip with a
   connective. Anthers produce pollen grains for pollination.
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  Plants and animals     4. Gynoecium (female reproductive part) - centrally located. It consists of a
                            collection of one or more carpels which organise to form one or more pistils.
                            Each pistil has three parts -
                               z    Ovary - It is the swollen basal part, one to many chambered (called
                                    locules) containing ovules which get fertilized to form seeds and the,
                                    fertilized ovary forms the fruit.
               Notes                                      Style
                                                                                Stigma
                                                                                             Petal
                                                                                             Stamens
Carpel
Sepal
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A. Variations in sepals and petals                                                             Plants and animals
(i) Polysepalous and Polypetalous (poly - free)- sepals or petals are free
      respectively.
(ii) Gamosepalous and Gamopetalous (gamo - united)- all sepals or petals are
      fused, respectively.
(iii) Perianth - Sepals and petals not distinguishable e.g. onion
B. Variations in Stamens (Fig. 7.23)                                                  Notes
     The stamens show variation in their cohesion (fusion).
       (i) Monadelphous - filaments fused into one bundle but anthers are free e.g.
           china rose
      (ii) Diadelphous – filaments fused to form two bundles e.g. pea
     (iii) Polyadelphous–filaments fused to form many bundles e.g., lemon
     (iv) Syngenecious – filaments are free but anthers are fused e.g. sunflower
      (v) Synandrous – stamens are fused throughout the length e.g., cocks-comb.
     Other variations in stamens are as follows
     (vi) Epipetalous – stamens are attached to petals by their filaments but anthers
           are free e.g., brinjal
    (vii) Didynamous – four stamens, two short and two long e.g. tulsi
   (viii) Tetradynamous – six stamens, inner four are long and outer two are short
           e.g., mustard
Notes
                                                         Do you know
                               Some plants like cashew nuts and mango have neuter, bisexual and
                                              unisexual flowers on the same tree.
                         7.3.2 Placentation
                         It is the manner in which placentae are distributed in the ovary. Placenta is the point
                         of attachment of ovules (or future seed) in the ovary.
                         Types of placentation (Fig. 7.26)
                         (i) Marginal - The ovary is monocarpellary and one chambered and ovules are
                                arranged along the fused margins of the single carpel. e.g. pea, gram.
                         (ii) Axile - Ovary is polycarpellary syncarpous, having many chambers and ovules
                                present on the placenta develop from the central axis formed by the fusion of
                                the margins of two or more carpels e.g. China rose, tomato, bhindi,
                         (iii) Parietal - Ovary is polycarpellary and syncarpous, having one chamber and
                                ovules are attached on its inner wall where margins of adjoining carpels meet
                                e.g, mustard, cucumber,
                         (iv) Basal – Ovary is bi-or polycarpellary, syncarpous, having one chamber and
                                placenta develops at the base of ovary and bears a single ovule e.g. sunflower.
                         (v) Free central – Ovary is syncarpous and polycarpellary but unilocular as septae
                                are absent. In the central part of the ovary the placenta bears many groups of
                                ovules e.g. Dianthus, Primula.
                                    Fig. 7.25 Types of placentation (a) Marginal, (b) Axile, (c) Parietal,
                                                 (d) Free central, (e) Basal, (f) Superficial
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(vi) Superficial - Ovary is polycarpellary syncarpous and multilocular in which                                                Plants and animals
     entire inner walls of chambers are lined with placental tissue so that ovules
     develop all around e.g.,water lily (Nymphaea)
 7.4 INFLORESCENCE
Inflorescence is the arrangement of flowers on the floral axis called peduncle.
:Inflorescence could be terminal or axillary.
7.4.1 Types of inflorescence
The various types of inflorescence depend upon the type of branching of peduncle
and arrangement of flowers. There are two major types of inflorescence
(i) Racemose. The main axis does not end in a -flower but continues to grow.
(ii) Cymose. The main axis ends in a flower and the growth is limited.
The major differences between the two are given in table 7.14
           Table 7.14 Differences between Racemose and Cymose
                                  inflorescence
                    Racemose                                                       cymose
 1. Main axis shows unlimited growth                               Growth is limited
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  Plants and animals                     Table 7.15 Types of Racemose Inflorescence
                                                         (Fig. 7.20)
2. Spike Like raceme but the flowers are sessile Achyranthes (‘Latzira’)
                         5. Spadix      Like spike but the axis is fleshy and enclosed   Colocassia, banana
                                        by a large showy bract (Spathe)
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                                                                                                  Notes
       Raceme                             Spikelet
                        Spike                                   Catkin               Spadix
                                          (b)                    (c)
                        (a)
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  Plants and animals 7.4.2 Special types of inflorescence
                     1. Hypanthodium (Fig.7.28a) - The fleshy receptacle forms a cup like cavity and
                         has an apical opening. The male and female flowers are borne on the inner wall
                         of the cavity e.g. Fig, Peepal
                     2. Cyathium (Fig. 7.28b) - A type of inflorescence characteristic of Euphorbia,
                         in which a cup shaped involucre encloses a single female flower surrounded by
                         a number of male flowers. A nectary is present at the rim of involucre,
               Notes
                     3. Verticillaster (Fig. 7.28c)- It is a series of condensed dichasial cyme at each
                         node with a cluster of sessile flowers in the axil of leaves e.g. Ocimum (Tulsa),
                         Salvia,
                                               Ostiole
                                                          Male flower
                                                                                             Male
                                                                                             flower
                                                             Female
                                                             flower
                          7.5 FRUIT
                         A true fruit is a ripened ovary that develops after fertilization. Ovules develop into
                         seeds and the ovary wall matures into fruit wall which is now called pericarp. The
                         pericarp may be thick or thin. In fleshy fruits like mango, pericarp is thick and
                         differentiated into three regions-(a) epicarp forms the skin of the fruit
                         (b) mesocarp, middle pulpy and (c) endocarp inner hard and stony (coconut,
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mango) or often thin membranes (orange). In dry fruits pericarp, is thin, dry, papery             Plants and animals
or thick and woody but not divided into three regions.
Sometimes along with ovary other floral parts like thalamus, receptacle or calyx
may develop as part of fruit, such fruits are-called false fruits. e.g. apple, pear
(thalamus), fig (receptacle).
Parthenocarpic fruit -It is a fruit that develops without fertilization. It is seedless
or has non-viable seeds e.g, banana, grapes. Horticulturists are producing such fruits Notes
artificially.
7.5.1 Kinds of fruits - There are three basic types
1. Simple fruit - Develops from single mono-to polycarpellary, syncarpous (fused)
    ovary e.g, pea, tomato.
2. Aggregate fruit - Collection (etaerio) of simple fruits or fruitlets on same
   thalamus developing from polycarpellary, apocarpous (free carpels) ovary e.g.
   Calotropis and Ranunculus.
3. Composite or multiple fruit - Fruit develops from a number of flowers
   juxtaposed together or from inflorescence e.g. mulberry, pineapple.
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  Plants and animals                         Table 7.18 Common Fruits and their edible parts.
                              Names                                 Types                                           Edible Parts
                          1. Banana                      Berry – simple, fleshy                              Mesocarp and endocarp
                          2. Apple                      Pome – simple, fleshy                                Fleshy thalamus
                          3. Coconut                     Fibrous Drupe – simple, fleshy                      Endosperm
                          4. Custard Apple              Etaerio of Berries – aggregate                       Pericarp
               Notes
                          5. Date Palm                   Berry – simple, fleshy                              Pericarp
                          6. Cashew Nut                  Nut – simple, dry indehiscent                       Peduncle and Cotyledons
                          7. Mango                      Drupe – simple, fleshy                               Mesocarp
                          B. Orange                      Hesperidium – simple, fleshy                        Juicy hairs from endocarp,
                          9. Tomato                     Berry – simple, fleshy                               Pericarp and Placentae
                         10. Pear                       Pome – simple, fleshy                                Fleshy thalamus
                         11. Pineapple                  Sorosis – composite                                  Outer portion of receptacle,
                                                                                                             bracts and perianth
                         12. Fig                        Syconous – composite                                 Fleshy receptacle
                         13. Litchi                     Nut – simple                                         Juicy aril
                         14. Wheat                       Caryopsis – simple dry indehiscent                  Starchy endosperm
                         15. Strawberry                  Etaerio of achenes- aggregate                       Succulent thalamus
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  Plants and animals     z   Stomata help in gaseous exchange and allow loss of water vapour during
                             transpiration.
                         z   Special structures like bulliform cells, hydathodes and hairs occur in leaves of
                             some plants.
                         z   Flower is a modified shoot.
                         z   A typical flower has accessory whorls i.e., calyx and corolla and reproductive
               Notes         or essential whorls i.e., androecium (male) and gynoecium (female).
                         z   Flowers may be bisexual, unisexual or neuter; actinomorphic or zygomorphic;
                             hypogynous, perigynous or epigynous.
                         z   Variations occur in floral parts.
                         z   Placentation is the manner in which placentae bearing ovules are distributed in
                             the ovary. It is of many types.
                         z   Inflorescence is the arrangement of flowers on the floral axis.
                         z   It has two major types - racemose and cymose.
                         z   Hypanthodium, verticillaster and cyathium are special types of inflorescence.
                         z   Fruit is a ripened ovary that develops after fertilization
                         z   Ovules develop into seeds and the ovary wall matures into fruit wall called the
                             pericarp which may be thin or differentiated into epicarp, mesocarp and
                             endocarp.
                         z   Fruits may be true or false and categorized into simple, aggregate or composite
                             types.
                         z   Simple fruits may be dry (dehiscent or indehiscent) or fleshy.
                         z   A fruit that develops without fertilization is called parthenocarpic fruit.
                                  TERMENAL EXERCISES
                         1. Differentiate between
                              (i) Dicot stem and monocot stem
                             (ii) Root and stem
                            (iii) Racemose and cymose inflorescence
                            (iv) Stoma and hydathode
                             (v) True fruit and false fruit
                            (vi) Dicot and monocot leaf
                         2. Explain the different types of underground modified stem?
                         3. Explain the process of secondary growth in dicot stem.
                         4. Draw and label the vertical section of dicot leaf.
                         5. Define the following
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     (a) Flower         (b) Actinomorphic (c) Heterophylly                        Plants and animals
     (d) Phyllotaxy     (e) Hypogynous         (f) Parthenocarpic fruit
     (g) Venation.
6. What is cork cambium? State its functions.
7 Draw labelled diagrams of the following
     (a) Raceme and corymb inflorescence
     (b) Axile and parietal placentation                                         Notes
8. What is a fruit? Enlist the various types of simple- fleshy fruits giving one
    example of each type.
9. What are the edible parts of the following fruits
     (a) Mango          (b) Orange             (c) Apple
     (d) Banana         (e) Coconut            (f) Cashew nut
10. Match the following of column A with that of column B
            A                                     B
     (a) Tendril                          (i) Protection
     (b) Stolon                          (ii) Food, storage
     (c) Thorn                          (iii) Reproduction
     (d) Tuber                          (iv) Photosynthesis
     (e) Capitulum                       (v) Climbing
      (f) Phylloclade                   (vi) Sunflower
11. Name the type of modification of an underground, non-green structure bearing
    nodes and internodes and ‘eyes’.
12. If a section of stem shows scattered vascular bundles which are closed, have
    ‘Y’ shaped xylem and are surrounded by bundle sheath; what group of plant
    is it?
13. What is the region ouside the phellogen known as?
14. When the cambium is less active which type of wood does it produce?
2. (a) - (v) (b) - (iv) (c) - (ii) (d) - (i) (e) - (iii)
     4. Axile                                                                              Notes
7.13 1. When the main axis ends in a flower and the growth is limited
4. Capitulum, Hypanthodium
5. (a) - (v) (b) - (iv) (c) - (ii) (d) - (iii) (e) - (i)
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